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Literatur und Schriften


Pseudocalotes
Spitzkopfagame

HALLERMANN, J. (2009): The genera Pseudocalotes and Bronchocela in south East Asia: Taxonomy and distribution. - Abstracts of presentations hold on DeAGAMIS the 1st International Symposium on Agamid Lizards. Bonner Zoologische Beiträge, Bonn, 56 (4): 301.

HALLERMANN, J. (2011): Neues bei den Spitzkopfagamen der Gattung Pseudocalotes. – Terraria, Münster, 6 (27): 10-12.

HALLERMANN, J. & W. BÖHME (2000): A review of the genus Pseudocalotes (Squamata: Agamidae), with description of a new species from West Malaysia. – Amphibia-Reptilia, 21: 193-210.

MANTHEY, U. & W. GROSSMANN (1997): Genus Pseudocalotes. - In: Amphibien und Reptilien Südostasiens. Natur und Tier – Verlag, Münster: 198-200.


Pseudocalotes andamanensis (BOULENGER, 1891)

Green Crestless Forest Lizard

BOULENGER, G.A. (1891): On new or little known Indian and Malayan reptiles and batrachians. - Ann. Mag. nat. Hist., (6) 8: 288-292.

HARIKRISHNAN & VASUDEVAN (2013): Rediscovery of Calotes andamanensis Boulenger, 1891, and assessment of its generic allocation (Squamata: Sauria: Agamidae). – Herpetozoa, Wien, 26 (1/2): 3-13.

ISHWAR, N.M. & I. DAS (1998): Rediscovery of Calotes andamanensis Boulenger 1891, and a reassessment of the type locality. - J. Bombay Nat. Hist. Soc., 95: 513-514.


Pseudocalotes austeniana (ANNANDALE, 1908)

Arbor Hills Agama

ANNANDALE, N. (1908): Description of a new species of lizard of the genus Salea from Assam. - Records of the Indian Museum, 2: 97.

DAS, A. & I. DAS (2007): Rediscovery of Mictopholis austeniana (Annandale, 1908) (Squamata: Agamidae). - Curr. Herpetol., 26 (1): 45-47.

MAHONY, S. (2010): Systematic and taxomonic revaluation of four little known Asian agamid species, Calotes kingdonwardi Smith, 1935, Japalura kaulbacki Smith, 1937, Salea kakhienensis Anderson, 1879 and the monotypic genus Mictopholis Smith, 1935 (Reptilia: Agamidae). – Zootaxa, 2514: 1-23.


Pseudocalotes baliomus HARVEY, SHANEY, HAMIDY, KURNIAWSAN & SMITH, 2017

Spot-Shouldered False Garden Lizard

HARVEY, M.B., SHANEY, K., HAMIDY, A., KURNIAWASN, N. & E.N. SMITH (2017): A new species of Pseudocalotes (Squamata: Agamidae) from the Bukit Barisan Range of Sumatra with an estimation of its phylogeny. – Zootaxa, 4276 (2): 215-232.

We describe a new species of Pseudocalotes from montane forests of the central, western coast of Sumatra. The combination of 3 or 4 interoculabials, slightly enlarged and heavily keeled scales on the lower flanks, a white scapular spot, a relatively long fifth toe, bicarinate lamellae at the base of Toe III, absence of a postrictal modified scale, and three rows of paravertebrals directed dorsally and posteriorly distinguishes the new species from its congeners on Sumatra and Java. Among the new characters defined in this study, counts of subdigital lamellae within the “span of Toe V” proved particularly useful for diagnosing species of Pseudocalotes. Phylogenetic analysis of DNA sequences identified a monophyletic group containing the Sumatran and Javan Pseudocalotes. Based on the phylogeny, we define a P. cybelidermus Group containing P. cybelidermus and P. guttalineatus and a P. tympanistriga Group containing P. tympanistriga, P. rhammanotus, and the new species. Combinations of five morphological characters define each of these two clades. As currently defined, Pseudocalotes is polyphyletic; Javan and Sumatran species are more closely related to other draconines such as Dendragama boulengeri than they are to some mainland species.


Pseudocalotes brevipes WERNER, 1904

Vietnam False Bloodsucker

HALLERMANN, J. (2000): The taxonomic status of Acanthosaura fruhstorferi Werner, 1904 and Calotes brevipes Werner, 1904 (Squmata: Agamidae). – Mitt. Mus. Naturk. Berlin, Zool. Reihe, 76 (1): 143-150.

WERNER, F. 1904: Beschreibung neuer Reptilien aus den Gattungen Acanthosaura, Calotes, Gastropholis und Typhlops. Zoologischer Anzeiger, 27: 461-464.


Pseudocalotes cybelidermus HARVEY, HAMIDY, KURNIAWAN, SHANEY & SMITH, 2014

Purple-throated False Garden Lizard

ANNANDALE, N. (1908): Description of a new species of lizard of the genus Salea from Assam. - Records of the Indian Museum, 2: 97.

HARVEY, M.B., HAMIDY, A., KURNIAWAN, N., SHANEY, K. & E.N. SMITH (2014): Three new species of Pseudocalotes (Squamata: Agamidae) from southern Sumatra, Indonesia. – Zootaxa, 3841 (2): 211–238.


Pseudocalotes dragon GRISMER, L.L., QUAH, E.S.H., WOOD, P.L. JR., ANUAR, S., MUIN, A., DAVIS, H.R., MURDOCH, GRISMER, J.L. COTA, M. & A.J. COBOS (2016)


Drogon’s False Garden Lizard

GRISMER, L.L., QUAH, E.S.H., WOOD, P.L. JR., ANUAR, S., MUIN, A., DAVIS, H.R., MURDOCH, GRISMER, J.L. COTA, M. & A.J. COBOS (2016): Dragons in the mist: three new species of Pseudocalotes Fitzinger (Squamata: Agamidae) from the sky island archipelago of Peninsular Malaysia - Zootaxa, 4136 (3): 461–490.

An integrative taxonomic analysis is used to delimit and describe three new species of Pseudocalotoes from the sky island archipelago of the Banjaran (=mountain range) Titiwangsa of Peninsular Malaysia. Pseudocalotes drogon sp. nov., from Fraser’s Hill, Pahang is basal to the sister species P. larutensis from Bukit Larut, Perak in the Banjaran Bintang and the new species P. rhaegal sp. nov. from Cameron Highlands, Pahang. Pseudocalotes drogon sp. nov. is differentiated from all other species of Psuedocalotes by having the combination of a flat rostrum; seven postrostrals; an interparietal; 11 cir-cumorbitals; five canthals; 7–10 superciliaries; one scale between the rostral and nasal; nine supralabials; eight infralabi-als; 10 postnasal-suborbital scales; four postmentals; five or six sublabials; five or six chinshields; 47 smooth, wide, gular scales; weak transverse gular and antehumeral folds; two enlarged scales between the ear and eye; enlarged upper and low-er posttemporals; a single enlarged supratympanic; no enlarged postrictals; three large scales bordering the dorsal margin of the ear opening; large pretympanic scales; eight scales in the nuchal crest not separated by a gap; enlarged vertebral scales extending to the tip of the tail; keeled and non-plate-like scales on flanks; 51 midbody scales; midventrals smaller than dorsals; 19 subdigital lamellae on the fourth finger; 23 subdigital lamellae on the fourth toe; preaxial scales on third toe enlarged and spinose; subdigital lamellae not unicarinate; HW/HL 0.52; HL/SVL 0.31; no elbow or knee patches; and a male dewlap color of lime-green bearing a central yellow spot. Pseudocalotes rhaegal sp. nov. is differentiated from all other Psuedocalotes by having the combination of a convex rostrum; 6–8 postrostrals; an interparietal; nine or 10 circu-morbitals; five canthals; 7–10 superciliaries; one or two scales between the rostral and nasal scales; eight or nine suprala-bials; seven or eight infralabials; 11 or 12 postnasal-suborbital scales; four postmentals; four or five chinshields; 40–45 smooth, wide, gular scales; no transverse gular fold; a weak antehumeral fold; three or four enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; no large scales bordering the upper margin of the ear opening or in the pretympanic region; 6–8 enlarged nuchal crest scales not separated by a gap; enlarged vertebral scales extending to the base of the tail; weakly keeled, non-plate-like scales on the flanks; 52– 58 midbody scales; midventrals smaller than dorsals; 19–21 subdigital lamellae on the fourth finger; 22–26 subdigital la-mellae on the fourth toe; preaxial scales on the third enlarged and rounded; subdigital lamellae not unicarinate; HW/HL 0.50–0.54; HL/SVL 0.28–0.30; no elbow or knee patches; and female dewlap color yellow bearing a purple base. The analyses also indicated that the new species, P. viserion sp. nov. from Genting Highlands, Pahang in the southern section of the Banjaran Titiwangsa is the sister species of P. flavigula from Cameron Highlands 121 km to the north and can be separated from all other species of Psuedocalotes by having the combination of three postrostrals; 10 circumorbitals; four or five canthals; 5–7 superciliaries; rostral and nasals in contact; supralabials contacting the nasal; six or seven suprala-bials; six or seven infralabials; two or three postmentals; 47 or 48 smooth, flat, gular scales; three chinshields; weak trans-verse gular and antehumeral folds; two enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; 7–9 nuchal crest scales lacking gaps and not extending beyond midbody; weakly keeled and plate-like scales on the flanks; 35–38 midbody scales; ventrals smaller than dorsals; 22 or 23 subdigital lamellae on the fourth finger; 26 or 27 subdigital lamellae on the fourth toe; preaxial scales on the third toe not modified; subdigital scales not unicarinate; HW/HL 0.62; no white marking below the eye; dewlap in males yel-low; and no elbow or knee patches. Pseudocalotes rhaegal sp. nov. most likely occurs in syntopy with P. flavigula in Ta-nah Rata at Cameron Highlands and its discovery adds to a growing body of literature detailing the recent descriptions of several new, upland, closely related, sympatric species in Peninsular Malaysia. Another new population referred to here as Pseudocalotes sp. nov. from the Hala-Bala Wildlife Sanctuary, Betong District, Yala Province, Thailand is discussed. The discovery and description of these three new Pseudocalotes from the upland regions of Peninsular Malaysia continues to underscore the remarkably high herpetological diversity and ecological complexity in this sky island archipelago that is still underestimated, unappreciated, and unprotected.


Pseudocalotes dringi ALLERMANN & BÖHME, 2000


HALLERMANN, J. & W. BÖHME (2000): A review of the genus Pseudocalotes (Squamata: Agamidae), with description of a new species from West Malaysia. – Amphibia-Reptilia, 21: 193-210.


Pseudocalotes flavigula SMITH, 1924

Gelbkehl-Spitzkopfagame / Malaya False Bloodsucker / Yello-throated Forest Agamid

SMITH, M.A. (1924): Two new lizards and a new tree frog from the Malay Peninsula. - J. Fed. Malay. St. Mus., Singapore, 11:183-186.


Pseudocalotes floweri BOULENGER, 1912

Flowers Spitzkopfagame / Thai False Bloodsucker / Flower´s Foirest Agamid

BOULENGER, G.A. (1912): A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore incl. the adjacent islands. Reptilia and Amphibia. - London (Taylor & Francis), xiii + 298 S.


Pseudocalotes guttalineatus HARVEY, HAMIDY, KURNIAWAN, SHANEY & SMITH, 2014

Dash-lined False Garden Lizard

HARVEY, M.B., HAMIDY, A., KURNIAWAN, N., SHANEY, K. & E.N. SMITH (2014): Three new species of Pseudocalotes (Squamata: Agamidae) from southern Sumatra, Indonesia. – Zootaxa, 3841 (2): 211–238.


Pseudocalotes kakhienensis (ANDERSON, 1879)

Birmesische Bergagame / Burmese Mountain Agamid

MAHONY, S. (2010): Systematic and taxomonic revaluation of four little known Asian agamid species, Calotes kingdonwardi Smith, 1935, Japalura kaulbacki Smith, 1937, Salea kakhienensis Anderson, 1879 and the monotypic genus Mictopholis Smith, 1935 (Reptilia: Agamidae). – Zootaxa, 2514: 1-23.


Pseudocalotes khaonanensis CHAN-ARD, COTA, MAKCHAI & LAOTEOW 2008

Spitzkopfagame

CHAN-ARD, T., COTA, M., MAKCHAI, S. & S. LAOTEOW (2008): A new species of the genus Pseudocalotes (Squamata: Agamidae) from peninsular Thailand. - The Thailand Natural History Museum Journal, 3 (1): 25-31.


Pseudocalotes kingdonwardi (SMITH, 1935)

Kingdonward's Bloodsucker

MAHONY, S. (2010): Systematic and taxonomic revaluation of four little known Asian agamid species, Calotes kingdonwardi SMITH, 1935, Japalura kaulbacki SMITH, 1937. Salea kakhienensis ANDERSON, 1879 and the montypic genus Mictopholis SMITH, 1935 (Reptilia: Agamidae). – Tootaxa, Auckland, 2514: 1-23.

SMITH, M.A. (1937): Description of a new species of agamid lizard from upper Burma. – Journal of the Bombay Natural History Society, 39 (4): 755.

Pseudocalotes kingdonwardi kingdonwardi (SMITH, 1935)

Pseudocalotes kingdonwardi bapoensis (YANG & SU, 1979)


Pseudocalotes larutensis HALLERMANN & MCGUIRE, 2001


Bukit Larfut False Garden Lizard

HALLERMANN, J. & J.A. MCGUIRE (2001): A new species of Pseudocalotes (Squamata: Agamidae) from Bukit Larut, West Malaysia. – Herpetologica, 57 (3):255-265.


Pseudocalotes microlepis BOULENGER, 1888

Braune Spitzkopfagame / Burmese False Bloodsucker / Small-scaled Forest Agamid

ZIEGLER, T., VU N.T., LE, K.Q., NGUYEN, Q.T., HALLERMANN, J., LE, V.K. & M.H. THACH (2006): Neue Verbreitungsnachweise einiger wenig bekannter vietnamesischer Amphibien und Reptilien. – Sauria, Berlin, 29 (2): 29-40.

We provide new distribution records for one Vietnamese rhacophorid frog (Theloderma corticale from Ha Giang: Du Gia), one agamid lizard (Pseudocalotes microlepis from Da Nang(Quang Nam: Ba Na), four scincids (Eumeces tamdaoensis from Ha Giang, Du Gia; Lygosoma quadrupes from Nam Dinh: Giao Thuy; Scincella doriae from Da Nang/Quang Nam: Ba Na; Tropidophorus hainanus from Nam Dinh: Giao Thuy), one colubrid snake (Pseudoxenodon bambusicola from Ha Giang: Du Gia) and one viperid snake (Protobothrops mucrosquamatus from Thua Thien-Hue: A Luoi, and from Ha Giang: Du Gia). In addition, charactertistic morphological features are given as well as natural history notes.


Pseudocalotes poilani BOURRET, 1939

Laotian False Bloodsucker


GRISMER, L.L., QUAH, E.S.H., WOOD, P.L. JR., ANUAR, S., MUIN, A., DAVIS, H.R., MURDOCH, GRISMER, J.L. COTA, M. & A.J. COBOS (2016): Dragons in the mist: three new species of Pseudocalotes Fitzinger (Squamata: Agamidae) from the sky island archipelago of Peninsular Malaysia - Zootaxa, 4136 (3): 461–490.

Pseudocalotes rhaegal GRISMER, QUAH, WOOD, ANUAR, MUIN, DAVIS, MURDOCH, GRISMER, COTA & COBOS 2016 


Rhaegal’s False Garden Lizard

An integrative taxonomic analysis is used to delimit and describe three new species of Pseudocalotoes from the sky island archipelago of the Banjaran (=mountain range) Titiwangsa of Peninsular Malaysia. Pseudocalotes drogon sp. nov., from Fraser’s Hill, Pahang is basal to the sister species P. larutensis from Bukit Larut, Perak in the Banjaran Bintang and the new species P. rhaegal sp. nov. from Cameron Highlands, Pahang. Pseudocalotes drogon sp. nov. is differentiated from all other species of Psuedocalotes by having the combination of a flat rostrum; seven postrostrals; an interparietal; 11 cir-cumorbitals; five canthals; 7–10 superciliaries; one scale between the rostral and nasal; nine supralabials; eight infralabi-als; 10 postnasal-suborbital scales; four postmentals; five or six sublabials; five or six chinshields; 47 smooth, wide, gular scales; weak transverse gular and antehumeral folds; two enlarged scales between the ear and eye; enlarged upper and low-er posttemporals; a single enlarged supratympanic; no enlarged postrictals; three large scales bordering the dorsal margin of the ear opening; large pretympanic scales; eight scales in the nuchal crest not separated by a gap; enlarged vertebral scales extending to the tip of the tail; keeled and non-plate-like scales on flanks; 51 midbody scales; midventrals smaller than dorsals; 19 subdigital lamellae on the fourth finger; 23 subdigital lamellae on the fourth toe; preaxial scales on third toe enlarged and spinose; subdigital lamellae not unicarinate; HW/HL 0.52; HL/SVL 0.31; no elbow or knee patches; and a male dewlap color of lime-green bearing a central yellow spot. Pseudocalotes rhaegal sp. nov. is differentiated from all other Psuedocalotes by having the combination of a convex rostrum; 6–8 postrostrals; an interparietal; nine or 10 circu-morbitals; five canthals; 7–10 superciliaries; one or two scales between the rostral and nasal scales; eight or nine suprala-bials; seven or eight infralabials; 11 or 12 postnasal-suborbital scales; four postmentals; four or five chinshields; 40–45 smooth, wide, gular scales; no transverse gular fold; a weak antehumeral fold; three or four enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; no large scales bordering the upper margin of the ear opening or in the pretympanic region; 6–8 enlarged nuchal crest scales not separated by a gap; enlarged vertebral scales extending to the base of the tail; weakly keeled, non-plate-like scales on the flanks; 52– 58 midbody scales; midventrals smaller than dorsals; 19–21 subdigital lamellae on the fourth finger; 22–26 subdigital la-mellae on the fourth toe; preaxial scales on the third enlarged and rounded; subdigital lamellae not unicarinate; HW/HL 0.50–0.54; HL/SVL 0.28–0.30; no elbow or knee patches; and female dewlap color yellow bearing a purple base. The analyses also indicated that the new species, P. viserion sp. nov. from Genting Highlands, Pahang in the southern section of the Banjaran Titiwangsa is the sister species of P. flavigula from Cameron Highlands 121 km to the north and can be separated from all other species of Psuedocalotes by having the combination of three postrostrals; 10 circumorbitals; four or five canthals; 5–7 superciliaries; rostral and nasals in contact; supralabials contacting the nasal; six or seven suprala-bials; six or seven infralabials; two or three postmentals; 47 or 48 smooth, flat, gular scales; three chinshields; weak trans-verse gular and antehumeral folds; two enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; 7–9 nuchal crest scales lacking gaps and not extending beyond midbody; weakly keeled and plate-like scales on the flanks; 35–38 midbody scales; ventrals smaller than dorsals; 22 or 23 subdigital lamellae on the fourth finger; 26 or 27 subdigital lamellae on the fourth toe; preaxial scales on the third toe not modified; subdigital scales not unicarinate; HW/HL 0.62; no white marking below the eye; dewlap in males yel-low; and no elbow or knee patches. Pseudocalotes rhaegal sp. nov. most likely occurs in syntopy with P. flavigula in Ta-nah Rata at Cameron Highlands and its discovery adds to a growing body of literature detailing the recent descriptions of several new, upland, closely related, sympatric species in Peninsular Malaysia. Another new population referred to here as Pseudocalotes sp. nov. from the Hala-Bala Wildlife Sanctuary, Betong District, Yala Province, Thailand is discussed. The discovery and description of these three new Pseudocalotes from the upland regions of Peninsular Malaysia continues to underscore the remarkably high herpetological diversity and ecological complexity in this sky island archipelago that is still underestimated, unappreciated, and unprotected.


Pseudocalotes rhammanotus HARVEY, HAMIDY, KURNIAWAN, SHANEY & SMITH, 2014


Stitched-back False Garden Lizard

HARVEY, M.B., HAMIDY, A., KURNIAWAN, N., SHANEY, K. & E.N. SMITH (2014): Three new species of Pseudocalotes (Squamata: Agamidae) from southern Sumatra, Indonesia. – Zootaxa, 3841 (2): 211–238.


Pseudocalotes saravacensis INGER & STUEBING,1994




INGER, R.F. & R.B. STUEBING (1994): First record of the lizard genus Pseudocalotes (Lacertilia: Agamidae) in Borneo, with description of a new species. - Raffl. Bull. Zool., Singapore, 42 (4): 961-965.


Pseudocalotes tympanistriga GRAY, 1831

Indonesian False Bloodsucker

HARVEY, M.B., SHANEY, K., SIDIK, I., KURNIAWAN, N. & E.N. SMITH (2017): Endemic dragons of Sumatra’s volcanoes: New species of Dendragama (Squamata: Agamidae) and status of Salea rosaceum Thominot. -  Herpetological Monographs 31 (1): 69–97.

We discovered new populations of Dendragama at the northern and southern ends of Sumatra. High genetic distances and concordance of multiple, apparently independent diagnostic characters support our descriptions of these two populations as new species. We define new characters of the sublabial, tympanic, dorsal crest, and dorsolateral crest scales. The three species of Dendragama undergo remarkable color change in response to time of day and stress. Females lay 2–4 ovoid eggs, reach sexual maturity at about 60 mm snout–vent length, and likely produce multiple clutches each year. We remove Salea rosaceum Thominot from the synonymy of Dendragama boulengeri and argue that the unique holotype of S. rosaceum is a specimen of Pseudocalotes tympanistriga with incorrect locality information.

MERTENS, R. (1954): Über die javanische Eidechse Dendragama fruhstorferi und die Gattung Dendragama. – Senckenbergiana, 34 (4-6): 185-186.


Pseudocalotes viserion GRISMER, QUAH, WOOD, ANUAR, MUIN, DAVIS, MURDOCH, GRISMER, COTA & COBOS 2016 


Viserion’s False Garden Lizard 

GRISMER, L.L., QUAH, E.S.H., WOOD, P.L. JR., ANUAR, S., MUIN, A., DAVIS, H.R., MURDOCH, GRISMER, J.L. COTA, M. & A.J. COBOS (2016): Dragons in the mist: three new species of Pseudocalotes Fitzinger (Squamata: Agamidae) from the sky island archipelago of Peninsular Malaysia - Zootaxa, 4136 (3): 461–490.

An integrative taxonomic analysis is used to delimit and describe three new species of Pseudocalotoes from the sky island archipelago of the Banjaran (=mountain range) Titiwangsa of Peninsular Malaysia. Pseudocalotes drogon sp. nov., from Fraser’s Hill, Pahang is basal to the sister species P. larutensis from Bukit Larut, Perak in the Banjaran Bintang and the new species P. rhaegal sp. nov. from Cameron Highlands, Pahang. Pseudocalotes drogon sp. nov. is differentiated from all other species of Psuedocalotes by having the combination of a flat rostrum; seven postrostrals; an interparietal; 11 cir-cumorbitals; five canthals; 7–10 superciliaries; one scale between the rostral and nasal; nine supralabials; eight infralabi-als; 10 postnasal-suborbital scales; four postmentals; five or six sublabials; five or six chinshields; 47 smooth, wide, gular scales; weak transverse gular and antehumeral folds; two enlarged scales between the ear and eye; enlarged upper and low-er posttemporals; a single enlarged supratympanic; no enlarged postrictals; three large scales bordering the dorsal margin of the ear opening; large pretympanic scales; eight scales in the nuchal crest not separated by a gap; enlarged vertebral scales extending to the tip of the tail; keeled and non-plate-like scales on flanks; 51 midbody scales; midventrals smaller than dorsals; 19 subdigital lamellae on the fourth finger; 23 subdigital lamellae on the fourth toe; preaxial scales on third toe enlarged and spinose; subdigital lamellae not unicarinate; HW/HL 0.52; HL/SVL 0.31; no elbow or knee patches; and a male dewlap color of lime-green bearing a central yellow spot. Pseudocalotes rhaegal sp. nov. is differentiated from all other Psuedocalotes by having the combination of a convex rostrum; 6–8 postrostrals; an interparietal; nine or 10 circu-morbitals; five canthals; 7–10 superciliaries; one or two scales between the rostral and nasal scales; eight or nine suprala-bials; seven or eight infralabials; 11 or 12 postnasal-suborbital scales; four postmentals; four or five chinshields; 40–45 smooth, wide, gular scales; no transverse gular fold; a weak antehumeral fold; three or four enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; no large scales bordering the upper margin of the ear opening or in the pretympanic region; 6–8 enlarged nuchal crest scales not separated by a gap; enlarged vertebral scales extending to the base of the tail; weakly keeled, non-plate-like scales on the flanks; 52– 58 midbody scales; midventrals smaller than dorsals; 19–21 subdigital lamellae on the fourth finger; 22–26 subdigital la-mellae on the fourth toe; preaxial scales on the third enlarged and rounded; subdigital lamellae not unicarinate; HW/HL 0.50–0.54; HL/SVL 0.28–0.30; no elbow or knee patches; and female dewlap color yellow bearing a purple base. The analyses also indicated that the new species, P. viserion sp. nov. from Genting Highlands, Pahang in the southern section of the Banjaran Titiwangsa is the sister species of P. flavigula from Cameron Highlands 121 km to the north and can be separated from all other species of Psuedocalotes by having the combination of three postrostrals; 10 circumorbitals; four or five canthals; 5–7 superciliaries; rostral and nasals in contact; supralabials contacting the nasal; six or seven suprala-bials; six or seven infralabials; two or three postmentals; 47 or 48 smooth, flat, gular scales; three chinshields; weak trans-verse gular and antehumeral folds; two enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; 7–9 nuchal crest scales lacking gaps and not extending beyond midbody; weakly keeled and plate-like scales on the flanks; 35–38 midbody scales; ventrals smaller than dorsals; 22 or 23 subdigital lamellae on the fourth finger; 26 or 27 subdigital lamellae on the fourth toe; preaxial scales on the third toe not modified; subdigital scales not unicarinate; HW/HL 0.62; no white marking below the eye; dewlap in males yel-low; and no elbow or knee patches. Pseudocalotes rhaegal sp. nov. most likely occurs in syntopy with P. flavigula in Ta-nah Rata at Cameron Highlands and its discovery adds to a growing body of literature detailing the recent descriptions of several new, upland, closely related, sympatric species in Peninsular Malaysia. Another new population referred to here as Pseudocalotes sp. nov. from the Hala-Bala Wildlife Sanctuary, Betong District, Yala Province, Thailand is discussed. The discovery and description of these three new Pseudocalotes from the upland regions of Peninsular Malaysia continues to underscore the remarkably high herpetological diversity and ecological complexity in this sky island archipelago that is still underestimated, unappreciated, and unprotected.


Pseudocalotes ziegleri GRAY, 1831

HALLERMANN, J., NGUYEN, Q.T., ORLOV, N.L. & A. ANANANJEVA (2010): A new species of the genus Pseudocalotes (Squamata: Agamidae) from Vietnam. – Russ. J. Herpetol., 17 (1): 31-40.

HALLERMANN, J. (2010): Zwei neue Agamenarten in den Gattungen Pseudocalotes und Bronchocela. – Terraria, Münster, 5 (23): 76-77.


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