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Literatur und Schriften


Acanthosaura GRAY, 1831

Nackenstachler

ANANJEVA, N.B., ORLOV, N.L. & S.A. KALYABINA-HAUF (2008): Species of Acanthosaura Gray, 1831 (Agamidae: Sauria, Reptilia)of Vietnam: Results of Molecular and Morphological Study. - Biology Bulletin, 35 (2): 178-186.

On the basis of morphological and molecular data, taxonomic diversity of the agamid genus Acanthosaura in Vietnam is discussed. This genus is represented in Vietnam by four species of two complexes, A. capra and A. lepidogaster. DNA analysis and the absence of reliable finds suggest to exclude A. crucigera and A. armata from the faunal list of Vietnam. The recently described species A. nataliae Orlov, Nguyen and Nguyen, 2006 of the A. capra complex is considered. Two species are recognized in the A. lepidogaster complex, one of which, A. coronata Günther, 1861, Smith (1935) regarded as a synonym of A. lepidogaster (Cuvier, 1829).

BOBROV, V.V. (2015): Lizards (Reptilia, Sauria) from Vietnam in the herpetological collection of the Zoological Museum of Moscow University. 2. Genus Acanthosaura (Family Agamidae). - Current Studies in Herpetology 15 (3/4): 146–152. (in Russisch)

Data on the Acanthosaura lizards of the Vietnam fauna, deposited in the herpetological collection of the Zoological Museum of the Moscow University, are given. The following data are provided for every sample: accession number, capture location, habitat, and collector. The places of capture are shown on the map of the country.

HARBIG, P. & M. MARONDE (1983): Die Gattung Acanthosaura. – Sauria, Berlin, 5 (1): 31-33.

KALYABINA-HAUF, S., ANANJEVA, N.B., JOGER, U., LENK, P., MURTHY, R.W., STUART, B.L., ORLOV, N.L. & W.M.C.T. HO (2004): Molecular phylogeny of the genus Acanthosaura (Agamidae). – Current Herpetology, Kyoto, 23 (1): 7-16.

MANTHEY, U. & W. GROSSMANN (1979): Genus Acanthosaura. – In: Amphibien und Reptilien Südostasiens. Natur und Tier – Verlag, Münster: 154-156.

STARK, T. (2011): Acanthosaura. – Terra, 47 (1): 31-33.

WOOD Jr., P.L. (2009): A preliminary phylogeny of the genus Acanthosaura Gray 1931, inferred from mitochondrial and nuclear genes. - Abstracts of presentations hold on DeAGAMIS the 1st International Symposium on Agamid Lizards. Bonner Zoologische Beiträge, Bonn, 56 (4): 301.

WOOD Jr., P.L., JACKMAN, T.R., BAUER, A.M., GRISMER, L.L., THIRAKHUPT, K., AOWPHOL, A., GRISMER, J.L., ONN, C.K. & N. AHMAD (2010): Biogeography and systematics of the genus Acanthosaura Gray 1931 (Squamata: Agamidae) inferred from mitochondrial and nuclear genes. Abstracts of the Second International Symposium on Agamid Lizards «DeAgamis2». - Current Studies in Herpetology, 10 (3/4): 158.


Acanthosaura armata GRAY, 1827

Armored Pricklenape

GOLDBERG, S.R., BURSEY, C.R. & L.L. GRISMER (2016): Acanthosaura armata (Peninsular Horned Tree Lizard) endoparasites. - Herpetological Review, 47 (3): 458.

GOLDBERG, S.R. & L.L. GRISMER (2014): Acanthosaura armata (peninsular horned tree lizard) reproduction. - Herpetological Review, 45 (1): 122-123.

SMITS, J. (2004): Acanthosaura armata. – Lacerta, 62 (1): 3-5.

STEIOF, C., MANTHEY, U. & W. DENZER (1991): Acanthosaura armata (HARDWICKE & GRAY). – Sauria Suppl. Berlin, 1991, 13 (1-4): 217-222. (02.188)

Inhalt:
Beschreibung, Geschlechtsunterschiede, Verbreitung, Klima, Biotop, Lebensweise und Verhalten, Natürliche Nahrung, Terrarienhaltung, Futter, Verhalten im Terrarium, Zucht, Aufzucht, Literatur.


Acanthosaura bintangensis WOOD, GRISMER, GRISMER, AHMAD, CHAN & BAUER, 2009


WOOD Jr., P.L., GRISMER, J.L., GRISMER L.L., AHMAD, N., ONN, C.K. & A.M. BAUER (2009): Two new montane species of Acanthosaura Gray, 1831 (Squamata: Agamidae) from Peninsular Malaysia. – Zootaxa, 2012: 28-46.

Two new montane species of the agamid genus Acanthosaura (Gray, 1831) from Peninsular Malaysia are described based on having unique color pattern and scale characteristics. Acanthosaura bintangensis sp. nov. from Bukit Larut, Perak most closely resembles A. titiwangsaensis sp. nov. from Fraser’s Hill and Cameron Highlands, Pahang but differs from it by having more subdigital lamellae on the fourth finger; the presence of a row of enlarged keeled suborbital scales beginning and terminating with an elliptical keeled scale; by having more scales surrounding the occipital spines; having more spots in the dorsal pattern; and by having a black gular region with a yellow medial stripe. Both species closely resemble A. crucigera but, can be separated from it by having higher numbers of subdigital lamellae on the fourth finger and toe; higher numbers of trasverse scales in the canthus rostralis-supraorbital ridge; higher numbers of scales bordering the rostral scale, and lower numbers of scales between the fifth canthals. The discovery of a two new agamid lizards from montane forests in well-known areas of Peninsular Malaysia underscores the importance of continued field work in these regions.

Acanthosaura brachypoda ANANJEVA, ORLOV, NGUYEN & RYABOV, 2011


ANANJEVA, N.B., ORLOV, N.L., NGUYEN, T.T. & S.A. RYABOV (2011): A new species of Acanthosaura (Agamidae, Sauria) from Northwest Vietnam. – Russian Journal of Herpetology, 18 (3): 195-202.

A new species of Acanthosaura (Gray, 1831) from Fan Si Pan Mount, Lao Cai Province, northern Vietnam is described based on having a unique combinations of morphometric and meristic external characters and very distinctive color pattern. Acanthosaura brachypoda sp.nov. from Fan Si Pan Mount, northwest Vietnam most closely resembles A. lepidogaster but differs from it by larger body length, shorter limbs, lower number of ventral scales, larger scales on the head and spotted color pattern on the brown background. It is the 10th known species of genus Acanthosaura in Southeast Asia and 5th species of this genus in Vietnam.


Acanthosaura capra GÜNTHER,1861


Green Pricklenape

ORLOV, N.L., NGUYEN, Q.T. & V.S. NGUYEN (2006): A new Acanthosaura allied to A. capra Günther, 1861 (Agamidae, Sauria) from central Vietnam and southern Laos. – Russian Journal of Herpetology, 13 (1): 61-76.

GÜNTHER, A. (1861): Second list of Siamese reptiles. - Proc. Zool. Soc. London, 1861: 187-189.

GÜNTHER, A. (1861): Second list of Siamese reptiles. - Ann. Mag. Nat. Hist. (3) 8: 266-268.

NIETEN, S. (2009): Acanthosaura capra – der grüne Nackenstachler. – Terralog 89: 3.

Acanthosaura cardamomensis WOOD, GRISMER, GRISMER, NEANG, CHAV & HOLDEN, 2010

Nackenstachler

BOBROV, V.V. (2013): Acanthosaura cardamomensis (Reptilia, Sauria, Agamidae) as a lizard species new for the Vietnam fauna. – Modern Herpetology, 13 (1/2): 64-66. (in Russisch)

GOLDBERG, S.R. & L.L. GRISMER (2014): Acanthosaura cardamomensis reproduction. – Herpetological Review, 45 (1): 123.

WOOD Jr., P.L., GRISMER, L.L., GRISMER J.L., NEANG, T., CHAV, T. & J. HOLDEN (2010): A new cryptic species of Acanthosaura Gray, 1831 (Squamata: Agamidae) from Thailand and Cambodia. – Zootaxa, 2488: 22-38.


Acanthosaura coronata BOULENGER 1885


GÜNTHER, A. (1861): Second list of Siamese reptiles. - Proc. Zool. Soc. London, 1861: 187-189.

GÜNTHER, A. (1861): Second list of Siamese reptiles. - Ann. Mag. Nat. Hist. (3) 8: 266-268.


Acanthosaura crucigera BOULENGER, 1885

Boulenger´s Pricklenape

HEIJNEN, G. (1990): De verzorging en de mislikte kweek van de stekelnekagame (Acanthosaura crucigera). – Lacerta, 48 (6): 178-183. (02.187)

Summary:
A. crucigera is an arboreal lizard from southeast Asia. They are bizar looking, with spines on head, neck and body. The overall colour is brown. On the neck they have a diamond shaped dark brown spot. Maximum length is 33 cm.
In 1985 a couple was obtained.The male died a few weeks later, taking no food at all. It was infected with worms. The female was pregnant and laid 12 eggs. After that she died. The eggs did not hatch.
In 1987 one male and two females were bought. They thrived well on a diet of insects, worms and new-born mice. Temperatures in the terrarium varied between 26 and 18°C. February 1988 a female died, probably due to social stress. In Summer the remaining couple was kept in an outdoor terrarium. Back inside they mated and in December 1988 eleven eggs were produced. Incubated on peat at 29°C none hatched. In March 1989 ten eggs were laid. Eight had been fertilized but none hatched (at 27°C).
At this moment the female lives in an indoor vivarium, while the male occipies the livingroom. Once put together they avoid each other as much as possible.

STETTLER, P.H. (1974): Sozialer Streß bei Acanthosaura crucigera BOULENGER, 1885. – Das Aquarium. 8 (66): 552-556. (02.186)

Inhalt:
Umwelt, Freihaltung im Wohnraum, Name und verwandte Art, Gestalt, Färbung und Geschlechtsunterschied, Biotop, Haltung, Ernährung, Kopulation und Eiablage, Verhalten, Schlußbemerkungen.

STETTLER, P.H. (1975): Sociale ´stress` bij Acanthosaura crucigera BOULENGER, 1885. – Het Aquarium, 46 (5): 130-134.

ULBER, E. & E. ULBER (1984): Bemerkenswerte Verhaltensweisen bei Acanthosaura crucigera. – Sauria, Berlin, 6 (2): 29-30.

WÖHRLE, J. (1996): Nachzucht des Nackenstachlers Acanthosaura crucigera. – elaphe, 4 (4): 21-22.

WÖHRLE, J. (1997): Nachzucht von Nackenstachlern. – Die Aquarien- und Terrarien-Zeitschrift, 50 (1): 34.



Acanthosaura lepidogaster CUVIER, 1829

Brown Pricklenape

HALLERMANN, J. (2000): The taxonomic status of Acanthosaura fruhstorferi Werner, 1904 and Calotes brevipes Werner, 1904 (Squmata: Agamidae). – Mitt. Mus. Naturk. Berlin, Zool. Reihe, 76 (1): 143-150.

MELL, R. (1952): Bodenrenner im subtropischen Bergwald. Der Nackenstachler (Gonocephalus lepidogaster). – Die Aquar. Terrar. Z., Stuttgart, 5 (6): 160-163.

WERNER, F. (1904): Beschreibung neuer Reptilien aus den Gattungen Acanthosaura, Calotes, Gastropholis und Typhlops. – Zool. Anz. 27: 461-464.


Acanthosaura murphy NGUYEN, DO, HOANG, NGUYEN, McCORMACK, NGUYEN, ORLOV, NGUYEN & NGUYEN, 2018


NGUYEN, L.T., DO, D.T., HOANG, H.V., NGUYEN, T.T., McCORMACK, T.E.M., NGUYEN, T.Q., ORLOV, N.L., NGUYEN, V.D. & S.N. NGUYEN (2018): A New Species of the Genus Acanthosaura Gray, 1831 (Reptilia: Agamidae) from Central Vietnam. – Russian Journal of Herpetology, 25 (4): 259-274.

We describe a new agamid species of the genus Acanthosaura from Khanh Hoa and Phu Yen provinces, central Vietnam based on morphological and molecular data. The new species, Acanthosaura murphyi sp. nov., most closely resembles the Indochinese horned lizard (A. capra) and the Natalia horned lizard (A. nataliae), however, it can be differentiated from other congeners by the following characters: a large-sized species (snout-vent length = 103.7 – 127.3 mm in adult males, 123.0 mm in the adult female); cylindrical spine above posterior margin of eye present; spine on occiput between tympanum and nuchal crest absent; two scale rows between rostral scute and nostril; small slightly keeled scales on flank randomly intermixed with keeled medium and large scales; nuchal crests strongly developed; dorsal crests developed and separate from nuchal crests by a diastema; coloration in life of body varies from gray in adult to brown or emerald green in subadult, and brown to whitish gray ventral skin; and tail brown or greenish with indistinct light bands. Genetically, uncorrected sequence divergences of COI between the new species and its closest species, A. capra, is 6.5%. To date, the new species has been found in evergreen forest between 84 – 1060 m in Phu Yen and Khanh Hoa provinces only.


Acanthosaura nataliae ORLOV, TRUANG & SANG, 2006


ORLOV, TRUONG & VAN SANG (2006): A new Acanthosaura allied to A. capra GÜNTHER, 1861 (Agamidae, Sauria) from central Vietnam and southern Laos. – Russian J. Herp., 13 (1): 62.


Acanthosaura phongdienensis NGUYEN, JINm VO, NGUYEN, ZHOU, CHE, MURPHY & ZHANG, 2019


NGUYEN, S.N., JIN, J.Q., VO, B.D., NGUYEN, L.T., ZHOU, W.W., CHE, J., MURPHY, R.W. & Y.P. ZHANG (2019): A new species of Acanthosaura Gray 1831 (Reptilia: Agamidae) from central Vietnam. – Zootaxa, 4612 (4): 555-565.

Herein, we describe the new agamid species Acanthosaura phongdienensis sp. nov. from central Vietnam based on morphological and genetic data. Males of the new medium-sized species have a snout-vent length of up to 77.4 mm and females up to 64.7 mm. In both genders, the tail is longer than the snout-vent length. A spine occurs on each side of the neck, the rostral scute is entire, and the nuchal and dorsal crests are continuous. A postorbital spine is about half as long as the diameter of the eye-ball, and small lateral scales intermix with large, keeled scales whose tips point backwards and upwards. The new species differs genetically from the closely related species A. brachypoda, A. coronata, and A. lepidogaster by uncorrected p-distances of 13.6%, 21.7%, and 12.8%, respectively. This is the 13th species of Acanthosaura and the eighth species in the genus from Vietnam.


Acanthosaura phuketensis PAUWELS, SUMONTHA, KUNYA, NITIKUL, SAMPHANTHAMIT, WOOD & GRISMER, 2015


PAUWELS, O.S.G., SUMONTHA, M., KUNYA, K., ANITIKUL, A. SAMPHANTHAMIT, P. WOOD, P.L. JR. & L.L. GRISMER (2015): Acanthosaura phuketensis (Squamata: Agamidae), a new long-horned tree agamid from southwestern Thailand – Zootaxa, 4020 (3): 473–494.

We describe a new lowland forest-dwelling species of the genus Acanthosaura from Phuket Island and the Phuket mountain range in southwestern Thailand. Acanthosaura phuketensis sp. nov., the 11th species in the genus, seems most closely related to A. crucigera from Myanmar and western Thailand and A. cardamomensis from the Cardamom Mountains, but can be differentiated from them by a combination of morphological and coloration characteristics. This new discovery stresses the importance of preserving the last forest patches remaining on Phuket Island, home to three other squamate endemics.


Acanthosaura titiwangsaensis Wood, Grismer, Grismer, Ahmad, Chan & Bauer, 2009


BAKER, N. (2016): Swimming behaviour of Acanthosaura titiwangsaensis at Fraser's Hill, Peninsular Malaysia. – SEAVR (South East Asia Vertebrate Records), 2016: 47-48.

WOOD Jr., P.L., GRISMER, J.L., GRISMER L.L., AHMAD, N., ONN, C.K. & A.M. BAUER (2009): Two new montane species of Acanthosaura Gray, 1831 (Squamata: Agamidae) from Peninsular Malaysia. – Zootaxa, 2012: 28-46.

Two new montane species of the agamid genus Acanthosaura (Gray, 1831) from Peninsular Malaysia are described based on having unique color pattern and scale characteristics. Acanthosaura bintangensis sp. nov. from Bukit Larut, Perak most closely resembles A. titiwangsaensis sp. nov. from Fraser’s Hill and Cameron Highlands, Pahang but differs from it by having more subdigital lamellae on the fourth finger; the presence of a row of enlarged keeled suborbital scales beginning and terminating with an elliptical keeled scale; by having more scales surrounding the occipital spines; having more spots in the dorsal pattern; and by having a black gular region with a yellow medial stripe. Both species closely resemble A. crucigera but, can be separated from it by having higher numbers of subdigital lamellae on the fourth finger and toe; higher numbers of trasverse scales in the canthus rostralis-supraorbital ridge; higher numbers of scales bordering the rostral scale, and lower numbers of scales between the fifth canthals. The discovery of a two new agamid lizards from montane forests in well-known areas of Peninsular Malaysia underscores the importance of continued field work in these regions.


Acanthosaura tongbiguanensis LIU & RAO, 2019


LIU, S. & D. RAO (2019): A new species of the genus Acanthosaura from Yunnan, China (Squamata, Agamidae). – ZooKeys, 888: 105-132.

A new species of Acanthosaura from Yunnan, China is described based on unique morphometric and meristic external characters and a very distinctive color pattern. The fourteenth species recorded of this genus, Acanthosaura tongbiguanensis sp. nov., was previously considered A. lepidogaster although it more closely resembles A. crucigera. It can be separated from all other species of the genus by having different numbers of subdigital lamellae on the fourth finger and toe, and a different shape of the black eye patch. The new species differs genetically from investigated congeners by percentage distance of 14.46% to 23.27% (cytochrome b gene).


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