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Literatur und Schriften
Spitzkopfagame
HALLERMANN, J. (2009): The genera Pseudocalotes and Bronchocela in south East Asia: Taxonomy and distribution. - Abstracts of presentations hold on DeAGAMIS the 1st International Symposium on Agamid Lizards. Bonner Zoologische Beiträge, Bonn, 56 (4): 301.
HALLERMANN, J. (2011): Neues bei den Spitzkopfagamen der Gattung Pseudocalotes. – Terraria, Münster, 6 (27): 10-12.
HALLERMANN, J. & W. BÖHME (2000): A review of the genus Pseudocalotes (Squamata: Agamidae), with description of a new species from West Malaysia. – Amphibia-Reptilia, 21: 193-210.
MANTHEY, U. & W. GROSSMANN (1997): Genus Pseudocalotes. - In: Amphibien und Reptilien Südostasiens. Natur und Tier – Verlag, Münster: 198-200.
| Pseudocalotes andamanensis (BOULENGER, 1891) |
Green Crestless Forest Lizard
BOULENGER, G.A. (1891): On new or little known Indian and Malayan reptiles and batrachians. - Ann. Mag. nat. Hist., (6) 8: 288-292.
HARIKRISHNAN & VASUDEVAN (2013): Rediscovery of Calotes andamanensis Boulenger, 1891, and assessment of its generic allocation (Squamata: Sauria: Agamidae). – Herpetozoa, Wien, 26 (1/2): 3-13.
ISHWAR, N.M. & I. DAS (1998): Rediscovery of Calotes andamanensis Boulenger 1891, and a reassessment of the type locality. - J. Bombay Nat. Hist. Soc., 95: 513-514.
| Pseudocalotes baliomus HARVEY, SHANEY, HAMIDY, KURNIAWSAN & SMITH, 2017 |
Spot-Shouldered False Garden Lizard
HARVEY, M.B., SHANEY, K., HAMIDY, A., KURNIAWASN, N. & E.N. SMITH (2017): A new species of Pseudocalotes (Squamata: Agamidae) from the Bukit Barisan Range of Sumatra with an estimation of its phylogeny. – Zootaxa, 4276 (2): 215-232.
We describe a new species of Pseudocalotes from montane forests of the central, western coast of Sumatra. The combination of 3 or 4 interoculabials, slightly enlarged and heavily keeled scales on the lower flanks, a white scapular spot, a relatively long fifth toe, bicarinate lamellae at the base of Toe III, absence of a postrictal modified scale, and three rows of paravertebrals directed dorsally and posteriorly distinguishes the new species from its congeners on Sumatra and Java. Among the new characters defined in this study, counts of subdigital lamellae within the “span of Toe V” proved particularly useful for diagnosing species of Pseudocalotes. Phylogenetic analysis of DNA sequences identified a monophyletic group containing the Sumatran and Javan Pseudocalotes. Based on the phylogeny, we define a P. cybelidermus Group containing P. cybelidermus and P. guttalineatus and a P. tympanistriga Group containing P. tympanistriga, P. rhammanotus, and the new species. Combinations of five morphological characters define each of these two clades. As currently defined, Pseudocalotes is polyphyletic; Javan and Sumatran species are more closely related to other draconines such as Dendragama boulengeri than they are to some mainland species.
| Pseudocalotes brevipes WERNER, 1904 |
Vietnam False Bloodsucker
HALLERMANN, J. (2000): The taxonomic status of Acanthosaura fruhstorferi Werner, 1904 and Calotes brevipes Werner, 1904 (Squmata: Agamidae). – Mitt. Mus. Naturk. Berlin, Zool. Reihe, 76 (1): 143-150.
WERNER, F. 1904: Beschreibung neuer Reptilien aus den Gattungen Acanthosaura, Calotes, Gastropholis und Typhlops. Zoologischer Anzeiger, 27: 461-464.
| Pseudocalotes cybelidermus HARVEY, HAMIDY, KURNIAWAN, SHANEY & SMITH, 2014 |
Purple-throated False Garden Lizard
ANNANDALE, N. (1908): Description of a new species of lizard of the genus Salea from Assam. - Records of the Indian Museum, 2: 97.
HARVEY, M.B., HAMIDY, A., KURNIAWAN, N., SHANEY, K. & E.N. SMITH (2014): Three new species of Pseudocalotes (Squamata: Agamidae) from southern Sumatra, Indonesia. – Zootaxa, 3841 (2): 211–238.
| Pseudocalotes drogon GRISMER, L.L., QUAH, E.S.H., WOOD, P.L. JR., ANUAR, S., MUIN, A., DAVIS, H.R., MURDOCH, GRISMER, J.L. COTA, M. & A.J. COBOS (2016) |
Drogon’s False Garden Lizard
GRISMER, L.L., QUAH, E.S.H., WOOD, P.L. JR., ANUAR, S., MUIN, A., DAVIS, H.R., MURDOCH, GRISMER, J.L. COTA, M. & A.J. COBOS (2016): Dragons in the mist: three new species of Pseudocalotes Fitzinger (Squamata: Agamidae) from the sky island archipelago of Peninsular Malaysia - Zootaxa, 4136 (3): 461–490.
An integrative taxonomic analysis is used to delimit and describe three new species of Pseudocalotoes from the sky island archipelago of the Banjaran (=mountain range) Titiwangsa of Peninsular Malaysia. Pseudocalotes drogon sp. nov., from Fraser’s Hill, Pahang is basal to the sister species P. larutensis from Bukit Larut, Perak in the Banjaran Bintang and the new species P. rhaegal sp. nov. from Cameron Highlands, Pahang. Pseudocalotes drogon sp. nov. is differentiated from all other species of Psuedocalotes by having the combination of a flat rostrum; seven postrostrals; an interparietal; 11 cir-cumorbitals; five canthals; 7–10 superciliaries; one scale between the rostral and nasal; nine supralabials; eight infralabi-als; 10 postnasal-suborbital scales; four postmentals; five or six sublabials; five or six chinshields; 47 smooth, wide, gular scales; weak transverse gular and antehumeral folds; two enlarged scales between the ear and eye; enlarged upper and low-er posttemporals; a single enlarged supratympanic; no enlarged postrictals; three large scales bordering the dorsal margin of the ear opening; large pretympanic scales; eight scales in the nuchal crest not separated by a gap; enlarged vertebral scales extending to the tip of the tail; keeled and non-plate-like scales on flanks; 51 midbody scales; midventrals smaller than dorsals; 19 subdigital lamellae on the fourth finger; 23 subdigital lamellae on the fourth toe; preaxial scales on third toe enlarged and spinose; subdigital lamellae not unicarinate; HW/HL 0.52; HL/SVL 0.31; no elbow or knee patches; and a male dewlap color of lime-green bearing a central yellow spot. Pseudocalotes rhaegal sp. nov. is differentiated from all other Psuedocalotes by having the combination of a convex rostrum; 6–8 postrostrals; an interparietal; nine or 10 circu-morbitals; five canthals; 7–10 superciliaries; one or two scales between the rostral and nasal scales; eight or nine suprala-bials; seven or eight infralabials; 11 or 12 postnasal-suborbital scales; four postmentals; four or five chinshields; 40–45 smooth, wide, gular scales; no transverse gular fold; a weak antehumeral fold; three or four enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; no large scales bordering the upper margin of the ear opening or in the pretympanic region; 6–8 enlarged nuchal crest scales not separated by a gap; enlarged vertebral scales extending to the base of the tail; weakly keeled, non-plate-like scales on the flanks; 52– 58 midbody scales; midventrals smaller than dorsals; 19–21 subdigital lamellae on the fourth finger; 22–26 subdigital la-mellae on the fourth toe; preaxial scales on the third enlarged and rounded; subdigital lamellae not unicarinate; HW/HL 0.50–0.54; HL/SVL 0.28–0.30; no elbow or knee patches; and female dewlap color yellow bearing a purple base. The analyses also indicated that the new species, P. viserion sp. nov. from Genting Highlands, Pahang in the southern section of the Banjaran Titiwangsa is the sister species of P. flavigula from Cameron Highlands 121 km to the north and can be separated from all other species of Psuedocalotes by having the combination of three postrostrals; 10 circumorbitals; four or five canthals; 5–7 superciliaries; rostral and nasals in contact; supralabials contacting the nasal; six or seven suprala-bials; six or seven infralabials; two or three postmentals; 47 or 48 smooth, flat, gular scales; three chinshields; weak trans-verse gular and antehumeral folds; two enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; 7–9 nuchal crest scales lacking gaps and not extending beyond midbody; weakly keeled and plate-like scales on the flanks; 35–38 midbody scales; ventrals smaller than dorsals; 22 or 23 subdigital lamellae on the fourth finger; 26 or 27 subdigital lamellae on the fourth toe; preaxial scales on the third toe not modified; subdigital scales not unicarinate; HW/HL 0.62; no white marking below the eye; dewlap in males yel-low; and no elbow or knee patches. Pseudocalotes rhaegal sp. nov. most likely occurs in syntopy with P. flavigula in Ta-nah Rata at Cameron Highlands and its discovery adds to a growing body of literature detailing the recent descriptions of several new, upland, closely related, sympatric species in Peninsular Malaysia. Another new population referred to here as Pseudocalotes sp. nov. from the Hala-Bala Wildlife Sanctuary, Betong District, Yala Province, Thailand is discussed. The discovery and description of these three new Pseudocalotes from the upland regions of Peninsular Malaysia continues to underscore the remarkably high herpetological diversity and ecological complexity in this sky island archipelago that is still underestimated, unappreciated, and unprotected.
| Pseudocalotes dringi ALLERMANN & BÖHME, 2000 |
HALLERMANN, J. & W. BÖHME (2000): A review of the genus Pseudocalotes (Squamata: Agamidae), with description of a new species from West Malaysia. – Amphibia-Reptilia, 21: 193-210.
| Pseudocalotes flavigula SMITH, 1924 |
Gelbkehl-Spitzkopfagame / Malaya False Bloodsucker / Yello-throated Forest Agamid
SMITH, M.A. (1924): Two new lizards and a new tree frog from the Malay Peninsula. - J. Fed. Malay. St. Mus., Singapore, 11:183-186.
| Pseudocalotes floweri BOULENGER, 1912 |
Flowers Spitzkopfagame / Thai False Bloodsucker / Flower´s Foirest Agamid
BOULENGER, G.A. (1912): A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore incl. the adjacent islands. Reptilia and Amphibia. - London (Taylor & Francis), xiii + 298 S.
BOULENGER, G.A. (1912): Description of Pseudocalotes floweri in “A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore incl. the adjacent islands. Reptilia and Amphibia”. Taylor & Francis, London.
| Pseudocalotes guttalineatus HARVEY, HAMIDY, KURNIAWAN, SHANEY & SMITH, 2014 |
Dash-lined False Garden Lizard
HARVEY, M.B., HAMIDY, A., KURNIAWAN, N., SHANEY, K. & E.N. SMITH (2014): Three new species of Pseudocalotes (Squamata: Agamidae) from southern Sumatra, Indonesia. – Zootaxa, 3841 (2): 211–238.
| Pseudocalotes jingpo XU, GONG, HOU, WENG, LIU, DENG, HU & PENG, 2024 |
XU, Y., GONG, Y., HOU, M., WENG, S., LIU, S., DENG, J., HU, J. & PENG, L. (2024): A New Species of the Genus Pseudocalotes (Squamata: Agamidae) from Southwest Yunnan, China. – MDPI, Animals 2024; 14(6):826. https://doi.org/10.3390/ani14060826
In this study, a new species of the genus Pseudocalotes is described from Yingjiang County, Dehong Dai and Jingpo Autonomous Prefecture, Yunnan Province, China, based on four female specimens. It can be distinguished from its congeners by the following combination of characters: (1) interoculabials 3 or 4; (2) canthals 5–7; (3) cicrcumorbitals 8–11; (4) 1 scale between rostral and nasal; (5) interparietal 1; (6) superciliaries 4–6; (7) supralabials 6–7, the 1st in contact with the nasal; (8) infralabials 6–8; (9) transverse gular fold and antehumeral fold present; (10) 2–3 enlarged scales between eye and ear; (11) nuchal crest single, consists of 3–5 erected spines; (12) dorsal crest row single, discontinuous and low, located between two keeled, parallel and enlarged scale rows; (13) enlarged postrictals absent; (14) scales around midbody 53–62, dorsal body scales heterogenous in size and shape; (15) midventrals smaller than dorsals; (16) subdigital scales on the 4th finger 20–26, and on the 4th toe 24–29; (17) dorsal background coloration light taupe with four irregular brown patches along the middle of dorsal; (18) inner lips wathet, tongue aurantiacus, throat bluish black. The population from Yingjiang County was nested within a highly supported lineage, formed a sister taxon with P. kakhienensis (SH 97/UFB 100) and according to the p-distance, the new species differed from its congeners by 14.5% to 35.2% for NADH dehydrogenase subunit 2 (ND2) and 15.5% to 25.0% for NADH dehydrogenase subunit 4 (ND4).
| Pseudocalotes kakhienensis (ANDERSON, 1879) |
Birmesische Bergagame / Burmese Mountain Agamid
MAHONY, S. (2010): Systematic and taxomonic revaluation of four little known Asian agamid species, Calotes kingdonwardi Smith, 1935, Japalura kaulbacki Smith, 1937, Salea kakhienensis Anderson, 1879 and the monotypic genus Mictopholis Smith, 1935 (Reptilia: Agamidae). – Zootaxa, 2514: 1-23.
| Pseudocalotes khaonanensis CHAN-ARD, COTA, MAKCHAI & LAOTEOW 2008 |
Spitzkopfagame
CHAN-ARD, T., COTA, M., MAKCHAI, S. & S. LAOTEOW (2008): A new species of the genus Pseudocalotes (Squamata: Agamidae) from peninsular Thailand. - The Thailand Natural History Museum Journal, 3 (1): 25-31.
| Pseudocalotes kingdonwardi (SMITH, 1935) |
Kingdonward's Bloodsucker
MAHONY, S. (2010): Systematic and taxonomic revaluation of four little known Asian agamid species, Calotes kingdonwardi SMITH, 1935, Japalura kaulbacki SMITH, 1937. Salea kakhienensis ANDERSON, 1879 and the montypic genus Mictopholis SMITH, 1935 (Reptilia: Agamidae). – Tootaxa, Auckland, 2514: 1-23.
SMITH, M.A. (1937): Description of a new species of agamid lizard from upper Burma. – Journal of the Bombay Natural History Society, 39 (4): 755.
Pseudocalotes kingdonwardi kingdonwardi (SMITH, 1935)
Pseudocalotes kingdonwardi bapoensis (YANG & SU, 1979)
| Pseudocalotes larutensis HALLERMANN & MCGUIRE, 2001 |
Bukit Larfut False Garden Lizard
HALLERMANN, J. & J.A. MCGUIRE (2001): A new species of Pseudocalotes (Squamata: Agamidae) from Bukit Larut, West Malaysia. – Herpetologica, 57 (3):255-265.
| Pseudocalotes microlepis BOULENGER, 1888 |
Braune Spitzkopfagame / Burmese False Bloodsucker / Small-scaled Forest Agamid
MAKCHAI, S., CHAN-ARD, T. & Y. RASSAMEE (2010): On the incubation of Pseudocalotes microlepis (Squamata, Agamidae) from northern Thailand. – Thai. Nat. Hist. Mus. J., 4 (2): 93-95.
Six eggs of Pseudocalotes microlepis were moved from the pitnest on the ground in evergreen forest on a hillside at 1,400 m.asl. in Doi Pui, Chiang Mai Province, Northern Thailand, to incubate in the laboratory. All eggs were white, and oval in shape. The average size of the eggs was 7.6 x 15.6 mm. After 60 days three hatchlings emerged. The average size at birth measured from snout to vent was 22.3 mm, and the tail was 44.3 mm.
PAUWELS, O.S.G. & K. KUNYA (2009): Geographic distribution: Pseudocalotes microlepis (Small-scaled Montane Forest Lizard). – Herpetol. Rev., 40 (3): 364.
RAJU, V., DAS, S., Bag, P. & S. RAHA (2020): An Agamid Specimen from Manipur (India) Formerly Thought to be Pseudocalotes microlepis (Boulenger, 1888) Seems to Be an Undescribed Species. – Russian Journal of Herpetolog, 27 (1): 1-4.
For several years, record of Pseudocalotes microlepis from Manipur (India) has been a matter of debate among Indian herpetologists. We studied the specimen in question and found that the specimen differs from Pseudocalotes microlepis and all other described Agamid lizards of North East India and adjacent Myanmar. On the basis of morphological characters, we assign it provisionally to genus Calotes and it seems to represent a hitherto undescribed species. However, we cannot currently describe it as the specimen is in rather bad condition and thus precludes the description of important characters.
YU, X., DU, Y., FANG, M., LI, H. & L. LIN (2018): The mitochondrial genome of Pseudocalotes microlepis (Squamata: Agamidae) and its phylogenetic position in agamids. – As. Herpetol. Res., 9 (1): 24-34.
We describe the complete mitochondrial genome of a small-scaled forest agamid (Pseudocalotes microlepis), which is 17 873 bp in size, containing 13 PCGs, 2 rRNAs, 22 tRNAs and non-coding regions. The mitogenome has a typical gene order among squamates. 13 PCGs include 2 start codons (ATG and ATA), 3 stop codons (TAG, TAA and AGG), and an incomplete stop codon (T-). Codon usage analyses showed that CUA-Leu1 and CGG-Arg are the most frequently and rarely used codon, respectively. All 22 tRNAs were predicted to form canonical cloverleaf secondary structures, except for two tRNAs (tRNACys and tRNASer (AGY)) lacking the dihydorouridine (DHU) arm. The large noncoding region (control region) is 2 687 bp long (28.3% T, 18.2% C, 42.3% A, and 11.2% G), with four different types of repeating sequences. The phylogenetic tree resulting from BEAST analyses based on concatenated 2 rRNAs and 13 PCGs in sequence revealed that the newly sequenced P. microlepis, where the genus Acanthosaura were aggregated. Together with Calotes versicolor, they constitute the subfamily Draconinae.
ZIEGLER, T., VU N.T., LE, K.Q., NGUYEN, Q.T., HALLERMANN, J., LE, V.K. & M.H. THACH (2006): Neue Verbreitungsnachweise einiger wenig bekannter vietnamesischer Amphibien und Reptilien. – Sauria, Berlin, 29 (2): 29-40.
We provide new distribution records for one Vietnamese rhacophorid frog (Theloderma corticale from Ha Giang: Du Gia), one agamid lizard (Pseudocalotes microlepis from Da Nang(Quang Nam: Ba Na), four scincids (Eumeces tamdaoensis from Ha Giang, Du Gia; Lygosoma quadrupes from Nam Dinh: Giao Thuy; Scincella doriae from Da Nang/Quang Nam: Ba Na; Tropidophorus hainanus from Nam Dinh: Giao Thuy), one colubrid snake (Pseudoxenodon bambusicola from Ha Giang: Du Gia) and one viperid snake (Protobothrops mucrosquamatus from Thua Thien-Hue: A Luoi, and from Ha Giang: Du Gia). In addition, charactertistic morphological features are given as well as natural history notes.
| Pseudocalotes poilani BOURRET, 1939 |
Laotian False Bloodsucker
BOURRET, R. (1939): Description of Pseudocalotes poilani in: “Notes herpétologiques sur l'Indochine française. XVIII. Reptiles et batraciens reçus au Laboratoire des Sciences Naturelle de l'Université au cours de l'année 1939. Descriptions de quatre espèces et d'une variété nouvelles”. - Bulletin Général de l'Instruction Publique. Hanoi. 19: 5-39.
| Pseudocalotes rhaegal GRISMER, QUAH, WOOD, ANUAR, MUIN, DAVIS, MURDOCH, GRISMER, COTA & COBOS 2016 |
Rhaegal’s False Garden Lizard
GRISMER, L.L., QUAH, E.S.H., WOOD, P.L. JR., ANUAR, S., MUIN, A., DAVIS, H.R., MURDOCH, GRISMER, J.L. COTA, M. & A.J. COBOS (2016): Dragons in the mist: three new species of Pseudocalotes Fitzinger (Squamata: Agamidae) from the sky island archipelago of Peninsular Malaysia - Zootaxa, 4136 (3): 461–490.
An integrative taxonomic analysis is used to delimit and describe three new species of Pseudocalotoes from the sky island archipelago of the Banjaran (=mountain range) Titiwangsa of Peninsular Malaysia. Pseudocalotes drogon sp. nov., from Fraser’s Hill, Pahang is basal to the sister species P. larutensis from Bukit Larut, Perak in the Banjaran Bintang and the new species P. rhaegal sp. nov. from Cameron Highlands, Pahang. Pseudocalotes drogon sp. nov. is differentiated from all other species of Psuedocalotes by having the combination of a flat rostrum; seven postrostrals; an interparietal; 11 cir-cumorbitals; five canthals; 7–10 superciliaries; one scale between the rostral and nasal; nine supralabials; eight infralabi-als; 10 postnasal-suborbital scales; four postmentals; five or six sublabials; five or six chinshields; 47 smooth, wide, gular scales; weak transverse gular and antehumeral folds; two enlarged scales between the ear and eye; enlarged upper and low-er posttemporals; a single enlarged supratympanic; no enlarged postrictals; three large scales bordering the dorsal margin of the ear opening; large pretympanic scales; eight scales in the nuchal crest not separated by a gap; enlarged vertebral scales extending to the tip of the tail; keeled and non-plate-like scales on flanks; 51 midbody scales; midventrals smaller than dorsals; 19 subdigital lamellae on the fourth finger; 23 subdigital lamellae on the fourth toe; preaxial scales on third toe enlarged and spinose; subdigital lamellae not unicarinate; HW/HL 0.52; HL/SVL 0.31; no elbow or knee patches; and a male dewlap color of lime-green bearing a central yellow spot. Pseudocalotes rhaegal sp. nov. is differentiated from all other Psuedocalotes by having the combination of a convex rostrum; 6–8 postrostrals; an interparietal; nine or 10 circu-morbitals; five canthals; 7–10 superciliaries; one or two scales between the rostral and nasal scales; eight or nine suprala-bials; seven or eight infralabials; 11 or 12 postnasal-suborbital scales; four postmentals; four or five chinshields; 40–45 smooth, wide, gular scales; no transverse gular fold; a weak antehumeral fold; three or four enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; no large scales bordering the upper margin of the ear opening or in the pretympanic region; 6–8 enlarged nuchal crest scales not separated by a gap; enlarged vertebral scales extending to the base of the tail; weakly keeled, non-plate-like scales on the flanks; 52– 58 midbody scales; midventrals smaller than dorsals; 19–21 subdigital lamellae on the fourth finger; 22–26 subdigital la-mellae on the fourth toe; preaxial scales on the third enlarged and rounded; subdigital lamellae not unicarinate; HW/HL 0.50–0.54; HL/SVL 0.28–0.30; no elbow or knee patches; and female dewlap color yellow bearing a purple base. The analyses also indicated that the new species, P. viserion sp. nov. from Genting Highlands, Pahang in the southern section of the Banjaran Titiwangsa is the sister species of P. flavigula from Cameron Highlands 121 km to the north and can be separated from all other species of Psuedocalotes by having the combination of three postrostrals; 10 circumorbitals; four or five canthals; 5–7 superciliaries; rostral and nasals in contact; supralabials contacting the nasal; six or seven suprala-bials; six or seven infralabials; two or three postmentals; 47 or 48 smooth, flat, gular scales; three chinshields; weak trans-verse gular and antehumeral folds; two enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; 7–9 nuchal crest scales lacking gaps and not extending beyond midbody; weakly keeled and plate-like scales on the flanks; 35–38 midbody scales; ventrals smaller than dorsals; 22 or 23 subdigital lamellae on the fourth finger; 26 or 27 subdigital lamellae on the fourth toe; preaxial scales on the third toe not modified; subdigital scales not unicarinate; HW/HL 0.62; no white marking below the eye; dewlap in males yel-low; and no elbow or knee patches. Pseudocalotes rhaegal sp. nov. most likely occurs in syntopy with P. flavigula in Ta-nah Rata at Cameron Highlands and its discovery adds to a growing body of literature detailing the recent descriptions of several new, upland, closely related, sympatric species in Peninsular Malaysia. Another new population referred to here as Pseudocalotes sp. nov. from the Hala-Bala Wildlife Sanctuary, Betong District, Yala Province, Thailand is discussed. The discovery and description of these three new Pseudocalotes from the upland regions of Peninsular Malaysia continues to underscore the remarkably high herpetological diversity and ecological complexity in this sky island archipelago that is still underestimated, unappreciated, and unprotected.
| Pseudocalotes rhammanotus HARVEY, HAMIDY, KURNIAWAN, SHANEY & SMITH, 2014 |
Stitched-back False Garden Lizard
HARVEY, M.B., HAMIDY, A., KURNIAWAN, N., SHANEY, K. & E.N. SMITH (2014): Three new species of Pseudocalotes (Squamata: Agamidae) from southern Sumatra, Indonesia. – Zootaxa, 3841 (2): 211–238.
| Pseudocalotes saravacensis INGER & STUEBING,1994 |
INGER, R.F. & R.B. STUEBING (1994): First record of the lizard genus Pseudocalotes (Lacertilia: Agamidae) in Borneo, with description of a new species. - Raffl. Bull. Zool., Singapore, 42 (4): 961-965.
| Pseudocalotes tympanistriga GRAY, 1831 |
Indonesian False Bloodsucker
HARVEY, M.B., SHANEY, K., SIDIK, I., KURNIAWAN, N. & E.N. SMITH (2017): Endemic dragons of Sumatra’s volcanoes: New species of Dendragama (Squamata: Agamidae) and status of Salea rosaceum Thominot. - Herpetological Monographs 31 (1): 69–97.
We discovered new populations of Dendragama at the northern and southern ends of Sumatra. High genetic distances and concordance of multiple, apparently independent diagnostic characters support our descriptions of these two populations as new species. We define new characters of the sublabial, tympanic, dorsal crest, and dorsolateral crest scales. The three species of Dendragama undergo remarkable color change in response to time of day and stress. Females lay 2–4 ovoid eggs, reach sexual maturity at about 60 mm snout–vent length, and likely produce multiple clutches each year. We remove Salea rosaceum Thominot from the synonymy of Dendragama boulengeri and argue that the unique holotype of S. rosaceum is a specimen of Pseudocalotes tympanistriga with incorrect locality information.
MERTENS, R. (1954): Über die javanische Eidechse Dendragama fruhstorferi und die Gattung Dendragama. – Senckenbergiana, 34 (4-6): 185-186.
| Pseudocalotes viserion GRISMER, QUAH, WOOD, ANUAR, MUIN, DAVIS, MURDOCH, GRISMER, COTA & COBOS 2016 |
Viserion’s False Garden Lizard
GRISMER, L.L., QUAH, E.S.H., WOOD, P.L. JR., ANUAR, S., MUIN, A., DAVIS, H.R., MURDOCH, GRISMER, J.L. COTA, M. & A.J. COBOS (2016): Dragons in the mist: three new species of Pseudocalotes Fitzinger (Squamata: Agamidae) from the sky island archipelago of Peninsular Malaysia - Zootaxa, 4136 (3): 461–490.
An integrative taxonomic analysis is used to delimit and describe three new species of Pseudocalotoes from the sky island archipelago of the Banjaran (=mountain range) Titiwangsa of Peninsular Malaysia. Pseudocalotes drogon sp. nov., from Fraser’s Hill, Pahang is basal to the sister species P. larutensis from Bukit Larut, Perak in the Banjaran Bintang and the new species P. rhaegal sp. nov. from Cameron Highlands, Pahang. Pseudocalotes drogon sp. nov. is differentiated from all other species of Psuedocalotes by having the combination of a flat rostrum; seven postrostrals; an interparietal; 11 cir-cumorbitals; five canthals; 7–10 superciliaries; one scale between the rostral and nasal; nine supralabials; eight infralabi-als; 10 postnasal-suborbital scales; four postmentals; five or six sublabials; five or six chinshields; 47 smooth, wide, gular scales; weak transverse gular and antehumeral folds; two enlarged scales between the ear and eye; enlarged upper and low-er posttemporals; a single enlarged supratympanic; no enlarged postrictals; three large scales bordering the dorsal margin of the ear opening; large pretympanic scales; eight scales in the nuchal crest not separated by a gap; enlarged vertebral scales extending to the tip of the tail; keeled and non-plate-like scales on flanks; 51 midbody scales; midventrals smaller than dorsals; 19 subdigital lamellae on the fourth finger; 23 subdigital lamellae on the fourth toe; preaxial scales on third toe enlarged and spinose; subdigital lamellae not unicarinate; HW/HL 0.52; HL/SVL 0.31; no elbow or knee patches; and a male dewlap color of lime-green bearing a central yellow spot. Pseudocalotes rhaegal sp. nov. is differentiated from all other Psuedocalotes by having the combination of a convex rostrum; 6–8 postrostrals; an interparietal; nine or 10 circu-morbitals; five canthals; 7–10 superciliaries; one or two scales between the rostral and nasal scales; eight or nine suprala-bials; seven or eight infralabials; 11 or 12 postnasal-suborbital scales; four postmentals; four or five chinshields; 40–45 smooth, wide, gular scales; no transverse gular fold; a weak antehumeral fold; three or four enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; no large scales bordering the upper margin of the ear opening or in the pretympanic region; 6–8 enlarged nuchal crest scales not separated by a gap; enlarged vertebral scales extending to the base of the tail; weakly keeled, non-plate-like scales on the flanks; 52– 58 midbody scales; midventrals smaller than dorsals; 19–21 subdigital lamellae on the fourth finger; 22–26 subdigital la-mellae on the fourth toe; preaxial scales on the third enlarged and rounded; subdigital lamellae not unicarinate; HW/HL 0.50–0.54; HL/SVL 0.28–0.30; no elbow or knee patches; and female dewlap color yellow bearing a purple base. The analyses also indicated that the new species, P. viserion sp. nov. from Genting Highlands, Pahang in the southern section of the Banjaran Titiwangsa is the sister species of P. flavigula from Cameron Highlands 121 km to the north and can be separated from all other species of Psuedocalotes by having the combination of three postrostrals; 10 circumorbitals; four or five canthals; 5–7 superciliaries; rostral and nasals in contact; supralabials contacting the nasal; six or seven suprala-bials; six or seven infralabials; two or three postmentals; 47 or 48 smooth, flat, gular scales; three chinshields; weak trans-verse gular and antehumeral folds; two enlarged scales between the ear and eye; an enlarged upper and lower posttemporal; an enlarged supratympanic; no enlarged postrictals; 7–9 nuchal crest scales lacking gaps and not extending beyond midbody; weakly keeled and plate-like scales on the flanks; 35–38 midbody scales; ventrals smaller than dorsals; 22 or 23 subdigital lamellae on the fourth finger; 26 or 27 subdigital lamellae on the fourth toe; preaxial scales on the third toe not modified; subdigital scales not unicarinate; HW/HL 0.62; no white marking below the eye; dewlap in males yel-low; and no elbow or knee patches. Pseudocalotes rhaegal sp. nov. most likely occurs in syntopy with P. flavigula in Ta-nah Rata at Cameron Highlands and its discovery adds to a growing body of literature detailing the recent descriptions of several new, upland, closely related, sympatric species in Peninsular Malaysia. Another new population referred to here as Pseudocalotes sp. nov. from the Hala-Bala Wildlife Sanctuary, Betong District, Yala Province, Thailand is discussed. The discovery and description of these three new Pseudocalotes from the upland regions of Peninsular Malaysia continues to underscore the remarkably high herpetological diversity and ecological complexity in this sky island archipelago that is still underestimated, unappreciated, and unprotected.
| Pseudocalotes ziegleri GRAY, 1831 |
HALLERMANN, J., NGUYEN, Q.T., ORLOV, N.L. & A. ANANANJEVA (2010): A new species of the genus Pseudocalotes (Squamata: Agamidae) from Vietnam. – Russ. J. Herpetol., 17 (1): 31-40.
HALLERMANN, J. (2010): Zwei neue Agamenarten in den Gattungen Pseudocalotes und Bronchocela. – Terraria, Münster, 5 (23): 76-77.
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