Literatur und Schriften
Gonocephalus (KAUP, 1825)
Abbotts Winkelkopfagame / Abbott’s Anglehead Lizard

Winkelkopfagamen

ANANJEVA, N.B. & T.N. MATVEYEVA-DUJSEBAYEVA (1996): Some evidence of Gonocephalus species complex divergence basing on skin sense organs morphology, Russian Journal of Herpetology, 3 (1): 82-88.

DIONG, C.H., LOW, M.-H., TAN, E.-C., YONG, H.-S., HIKIDA, T. & H. OTA (2000): On the monophyly of the Agamid genus Gonocephalus Kaup, 1825 (Reptilia: Squamata) – A Chromosomal Perspective. - Current Herpetology, 19: 71–79.

EVERS, M. (2010): Winkelkopfagamen. – Natur und Tier-Verlag, Münster. 64 S.

EVERS, M. (2013): Der Traum vom Urwald im Wohnzimmer – Winkelkopfagamen (Gonocephalus) im Terrarium. – Reptilia, Münster, 18 (4): 50-59.

HONDA, M., OTA, H., SENGOKU, S., YONG, H.-S. & T. HIKIDA (2002): Molecular evaluation of phylogenetic significances in the divergent karyotypes of the genus Gonocephalus (Reptilia: Agamidae) from tropical Asia. – Zool. Sci., 19: 129-133.

MANTHEY, U. & W. DENZER (1991): Die Echten Winkelkopfagamen der Gattung Gonocephalus Kaup (Sauria: Agamidae) II. Allgemeine Angaben zur Biologie und Terraristik. – Sauria, Berlin, 13 (2): 19-22.

MANTHEY, U. & W. GROSSMANN (1979): Genus Gonocephalus - In: Amphibien und Reptilien Südostasiens. - Natur und Tier – Verlag, Münster: 177-191.

OTA, H., MATSUI, M., HIKIDA, T. & A. MORI (1992): Extreme karyotypic divergence between species of the genus Gonocephalus (Reptilia: Squamata: Agamidae) from Borneo and Australia. – Herpetologica, 48 (1): 120-124

Gonocephalus grandis and G. miotympanum have karyotypes consisting of 2N = 42 chromosomes forming two size groups. The smaller group comprises eight metacentric and 12 telocentric microchromosomes, whereas the larger group has 22 metacentric and submetacentric chromosomes. Slight interspecific differences were recognized in the centromeric positions of a few large pairs. These species are karyologically strikingly different from any other agamids including G. spinipes from Australia, and we suggest the possible presence of a zoogeographic dichotomy between Asian and Australian species currently assigned to the genus Gonocephalus.

SCHLEGEL, H. (1851): Descriptions de plusieurs expèces nouvelles du genre Lophyrus. – Bijdragen tot de Dierkunde 3 (1): 4-6.

WELTON, L.J., SILER, C.D., GRISMER, L.L., DIESMOS, A.C., SITES, J.W. & R.M. BROWN (2017): Archipelago-wide survey of Philippine forest dragons (Agamidae: Gonocephalus): multilocus phylogeny uncovers unprecedented levels of genetic diversity in a biodiversity hotspot. – Biol. J. Linn. Soc., 120: 410-426.

We utilize robust geographical genetic sampling, and phylogenetic analysis of a new multilocus dataset to provide the first inference of relationships among Philippine Gonocephalus, combined with estimates of putative species diversity, in this almost unknown island radiation. Our results reveal startling levels of undocumented diversity, genetically partitioned at a number of geographical levels across the archipelago. We present the first survey of genetic lineage diversity, coupled with an archipelago-wide clarification of geographical structure in a unique archipelago-endemic radiation. Philippine Gonocephalus have previously escaped the attention of biogeographers as a result of the taxonomic confusion associated with low numbers of preserved specimens in museum collections. With new vouchered material and genetic sampling from a comprehensive, archipelago-wide vertebrate biodiversity inventory, our findings join many recent studies in highlighting the unprecedented faunal diversity in one of the world’s most unique biodiversity conservation hotspots.




Gonocephalus bellii DUMÉRIL & BIBRON, 1837

Blaukehl-Winkelkopfagame / Bell´s Forest Dragon / Bell´s Anglehead Lizard

EVERS, M. (2007): Bizarre Regenwaldbewohner – Winkelkopfagamen. – Die Aquarien- und Terrarien-Zeitschrift, 60 (10): 70-75.

EVERS, M. (2010): Winkelkopfagamen. Gonocephalus bellii, G. chamaeleontinus, G. doriae abbotti. – Art für Art, Agamen 9, Münster (Natur und Tier-Verlag). 64 S.

KOBER, I. (2003): Erfahrungen mit der Blaukehl-Winkelkopfagame (Gonocephalus bellii) im Terrarium. – Draco, Münster, 4 (2): 80-84.
Inhalt:
Beschreibung, Eingewöhnung, Unterbringung und technische Ausstattung, Pflege und Verhalten im Terrarium, Bislang keine Nachzucht.

MANTHEY, U. & W. DENZER (1992): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). V. Die bellii-Gruppe. – Sauria, Berlin, 14 (3): 7-20.



Gonocephalus beyschlagi BOETTGER, 1892

Sumatra Forest Dragon

MANTHEY, U. & W. DENZER (1992): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). V. Die bellii-Gruppe. – Sauria, Berlin, 14 (3): 7-20.



Gonocephalus bornensis SCHLEGEL, 1848

Winkelkopfagame / Borneo Forest Dragon

MANTHEY, U. (1991): Gonocephalus denzeri sp.n. (Sauria: Agamidae) aus Sarawak, Borneo. - Salamandra, Bonn, 27 (3): 152-157. (00.076)

Summary:
Description of a new species of Gonocephalus from Borneo as Gonocephalus denzeri sp.n., closely related to G. liogaster. G. belli and G. miotympanum. Differs from all these species by no enlarged scales above ands below the tympanum and the enlarged scales in groups on the flanks.

MANTHEY, U. & W. DENZER (1992): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). V. Die bellii-Gruppe. – Sauria, Berlin, 14 (3): 7-20.

SCHLEGEL, H. (1851): Descriptions de plusieurs expèces nouvelles du genre Lophyrus. – Bijdragen tot de Dierkunde 3 (1): 4-6.




Gonocephalus chamaeleontinus LAURENTI, 1768

Chamäleon-Winkelkopfagame / Chameleon Forest Dragon / Chameleon Anglehead Lizard

EVERS, M. (2009): Ein bizarrer Kletterkünstler des tropischen Regenwaldes (I). – Die Aquarien- und Terrarienzeitschrift, 6 (10): 74-78.

EVERS, M. (2009): Ein bizarrer Kletterkünstler des tropischen Regenwaldes (II). – Die Aquarien- und Terrarienzeitschrift, 6 (11): 68-71.

EVERS, M. (2010): Winkelkopfagamen. Gonocephalus bellii, G. chamaeleontinus, G. doriae abbotti. – Art für Art, Agamen 9, Münster (Natur und Tier-Verlag). 64 S.

MANTHEY, U. & W. DENZER (1993): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). VI. Die chamaeleontinus-Gruppe. - Sauria, Berlin, 15 (2): 23-38. (1249)

SCHLEGEL, H. (1851): Descriptions de plusieurs expèces nouvelles du genre Lophyrus. – Bijdragen tot de Dierkunde 3 (1): 4-6.



Gonocephalus doriae (PETERS, 1871)

Dorias Winkelkopfagame / Peter´s Forest Dragon / Doria´s Anglehead Lizard

MANTHEY, U. & W. DENZER (1993): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). VI. Die chamaeleontinus-Gruppe. - Sauria, Berlin, 15 (2): 23-38.

PETERS, W. (1871): Uber neue Reptilien aus Ostafrika und Sarawak (Borneo), vorzüglich aus der Sammlung des Hrn. Marquis J. Doria zu Genua. - Mber. k. preuss. Akad. Wiss., Berlin [1871]: 566-581.


Gonocephalus doriae doriae (PETERS, 1871)


Gonocephalus doriae abbotti COCHRAN 1922

COCHRAN, D. M. 1922. Description of a new species of agamid lizard from the Malay Peninsula. Proc. US Natl. Mus. 60: 1-3.

DENZER, W., MANTHEY, U., BRADY, M.L. & O. JOHNSON (2020): Erstnachweis von Gonocephalus cf. abbotti (COCHRAN, 1922) auf Sumatra. – Sauria, Berlin, 42 (4): 49-56.

EVERS, M. (2010): Winkelkopfagamen. Gonocephalus bellii, G. chamaeleontinus, G. doriae abbotti. – Art für Art, Agamen 9, Münster (Natur und Tier-Verlag). 64 S.

GOLDBERG, S.R. & L.L. GRISMER (2014): Gonocephalus abbotti (Abbott's anglehead lizard) reproduction. - Herpetological Review, 45 (4): 696.

GROSSMANN, W. (2004): Das Portrait. Gonocephalus doriae abbotti COCHRAN. – Sauria, Berlin, 26 (3): 1-2.

Inhalt:
Verbreitung und Biotop / Systematische Einordnung und Beschreibung / Literatur.

HÖLßIG, C. (2012): Zur Haltung und Vermehrung von Gonocephalus doriae abbotti COCHRAN, 1922.- Sauria, Berlin, 34 (1): 43-50.

MANTHEY, U. & W. DENZER (1993): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). VI. Die chamaeleontinus-Gruppe. - Sauria, Berlin, 15 (2): 23-38.


Gonocephalus doriae brevis HARVEY et al. 2023

HARVEY, MICHAEL B., SARKER, GOUTAM C., SIDIK, IRVAN, KURNIAWAN, NIA, AND SMITH, ERIC N. 2023. A Contribution to the Systematics of Sunda Shelf Angle- Headed Dragons (Agamidae: Gonocephalus) with the Description of New Taxa from Sumatra. Herpetological Monographs, 37(1): 1-40



Gonocephalus grandis GRAY, 1845

Grosse Winkelkopfagame / Giant Forest Dragon / Great Anglehead Lizard

BAKER, N. (2016): Records of two lizards from riparian habitat at Sungai Salu, Perak, Peninsular Malaysia. . – SEAVR (South East Asia Vertebrate Records) 2016: 82-84.

DIECKMANN, M. (2013): Gonocephalus grandis (GRAY, 1845). – Iguana, 26 (1): 17-21.

GERICKE, F. (1984): Gonocephalus grandis. Erfolgreiche Haltung und Nachzucht. – Sauria, Berlin, 6 (4): 5-6.

GOLDBERG, S.R. & L.L. GRISMER (2014): Gonocephalus grandis (giant forest dragon). Reproduction. - Herpetological Review, 45 (4): 696.

HENNIG, U. & A.S. HENNIG (2003): Haltung und Nachzucht der Großen Winkelkopfagame. – Die Aquarien- und Terrarienzitschrift, 56 (1): 6-63.

LANDRY GREEN, M.E. (2019): Dissection of a Gonocephalus grandis (Gray, 1845): identifying cause of death in a rare captive lizard. – Captive & Field Herpetology, 3 (1): 10-18.

MANTHEY, U. & W. DENZER (1991): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). III. Gonocephalus grandis (GRAY, 1845). – Sauria, Berlin, 13 (3): 3-10.

Zusammenfassung:
Gonocephalus grandis steht morphologisach nicht in beziehung zu einer anderen Art der Gattung. Biologie und Verhalten von G. grandis werden im einzelnen beschrieben und eine Bibliographie mit Kurzinhaltsangaben vorgelegt.

MANTHEY, U. & W. DENZER (1991): The True Anglehead Agamas of the genus Gonocephalus KAUP (Sauria: Agamidae). III. Gonocephalus grandis (GRAY, 1825). - Sauria (E), Kempton Park, 1 (1): 3-9. (1151)

SHAHRIZA, S. (2018): Defense behavior of Gonocephalus grandis and Aphaniotis fusca (Squamata: Agamidae). – Phyllomedusa, 17 (1): 153-156.


Gonocephalus inauris HARVEY, SARKAR, SIDIK, KURNIAWAN & SMITH, 2023

HARVEY, MICHAEL B., SARKER, GOUTAM C., SIDIK, IRVAN, KURNIAWAN, NIA, AND SMITH, ERIC N. (2023): A Contribution to the Systematics of Sunda Shelf Angle- Headed Dragons (Agamidae: Gonocephalus) with the Description of New Taxa from Sumatra. Herpetological Monographs, 37(1): 1-40.

We review morphology of Sunda Shelf Gonocephalus with an emphasis on Sumatran and Javan species. At least 15 species and subspecies inhabit Peninsular Malaysia, Sumatra, Java, Borneo, and adjacent smaller continental islands. Following analysis of external morphology, we provide a dichotomous key to Sunda Shelf Gonocephalus and resolve two taxonomic problems with this group of lizards. Three populations of Gonocephalus doriae on Borneo, Peninsular Malaysia, and Sumatra are recognized as subspecies, because they lack concordance of multiple morphological differences and have low genetic divergence in a 556 base-pair fragment of the 16S rRNA gene. Described as new herein, G. doriae brevis from Aceh and North Sumatra differs from G. d. abbotti and G. d. doriae in usually having more scales around midbody and a relatively shorter tail with fewer dark bands. Though previously reported as lost, a syntype of G. d. doriae (MSNG 29152) is designated as lectotype, illustrated, and described. Also described as new, G. inauris from high elevations of the Bukit Barisan Range of Bengkulu, Sumatra, is a species of the G. megalepis Group differing from all congeners in having 7/6 (vs. 8–19) loreals separating the last canthal and supralabials, 8/8 (11–27) infraorbitals, and 58 (73–153) scales around midbody. In this new species, distinctly enlarged suboculars broadly contact its supralabials, whereas 1–4 lorilabials separate the suboculars and supralabials in congeners. High genetic divergence in the new species mirrors its high level of morphological divergence.



Gonocephalus interruptus BOULENGER,1885

Boulenger´s Forest Dragon

BOULENGER, G.A. (1885): Catalogue of the lizards on the British Museum (Natural History). I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. – London (Taylor & Francis). 436 S.



Gonocephalus klossi BOULENGER,1920

Kloss´ Forest Dragon

BOULENGER, G.A. (1920): Results of an expedition to Korinchi Peak 12.400 ft, Sumatra. III. Reptiles & batrachians collected in Korinchi, West Sumatra by Messrs. H.C. Robinson & C. Boden Kloss. J. - Federated Malay State Museum, Singapore, 8: 285-296.
Gonocephalus kuhlii SCHLEGEL, 1848

Winkelkopfagame / Forest Dragon

MANTHEY, U. & W. DENZER (1993): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). VI. Die chamaeleontinus-Gruppe. - Sauria, Berlin, 15 (2): 23-38.

SCHLEGEL, H. (1848): Descriptions de plusieurs expèces nouvelles du genre Lophyrus. - Bijdr. Dierk. (Amsterdam), 1: 4-6.



Gonocephalus lacunosus MANTHEY & DENZER,1991

Manthey´s Forest Dragon

MANTHEY, U. & W. DENZER (1991): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). I. Die megalepis-Gruppe mit Gonocephalus lacunosus sp. n. aus Nord-Sumatra. - Sauria, Berlin, 13 (1): 3-10. (1444)



Gonocephalus liogaster GÜNTHER,1872

Blauaugen-Winkelkopfagame / Tropical Forest Dragon / Blue-eyed Anglehead Lizard

GÜNTHER, A. (1872): On the reptiles and amphibians of Borneo. - Proc. Zool. Soc. London, 1872: 586-600.

MANTHEY, U. & W. DENZER (1992): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). V. Die bellii-Gruppe. – Sauria, Berlin, 14 (3): 7-20.



Gonocephalus megalepis BLEEKER,1860

Bleeker´s Forest Dragon

BLEEKER, P. (1860): Description of Gonocephalus megalepis. - In: “Reptilien van Agam”. – Nat. Tijdschr. Ned. Indie, 20: 325-329.

MANTHEY,U. & W. DENZER (1991): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae). I. Die megalepis-Gruppe mit Gonocephalus lacunosus sp. n. aus Nord-Sumatra. - Sauria, Berlin, 13 (1): 3-10.



Gonocephalus mjobergi SMITH, 1925

Winkelkopfagame / Forest Dragon

DENZER, W. & U. MANTHEY (2009): Remarks on the type specimen of Gonocephalus mjobergi Smith, 1925 (Sauria: Agamidae). – Bonner zoologische Beiträge, Bonn, 56 (4): 255-258.

Abstract:
The type specimen of Gonocephalus mjobergi Smith, 1925 has been re-examined. A remarkable charcter, longitudinal folds in the gular region has been discovered which is unique among agamids of the genus Gonocephalus Kaup. G. mjobergi has been compared to members of the genera Ptyctolaemus and Mantheyus and it is concluded that these genera form a monophyletic group and G. mjobergi is referred to as “Gen. A.” mjobergi until further material becomes available.

MANTHEY, U. & W. DENZER (1992): Die Echten Winkelkopfagamen der Gattung Gonocephalus KAUP (Sauria: Agamidae) IV Gonocephalus mjoebergi SMITH, 1925 und Gonocephalus robinsonii BOULENGER, 1908. – Sauria, Berlin, 14 (1): 15-19.

SMITH, M.A. (1925): On a collection of reptiles and amphibians from Mt. Murud, Borneo. - Sarawak Mus. J., Kuching, 8: 5-14.




Gonocephalus pyrius HARVEY, RECH, RIYANTO, KURNIAWAN & SMITH, 2021

Angle-headed Dragon

HARVEY, M.B., RECH, I., RIYANTO, A., KURNIAWAN, N. & E.N. SMITH (2021): A New Angle-Headed Dragon (Agamidae: Gonocephalus) from Montane Forests of the Southern Bukit Barisan Range of Sumatra. - Herpetologica 77 (1): 94-109.

We describe a new species of the Gonocephalus megalepis group and provide new morphological and distribution data for the three other species of the group. The combination of a wide gap between the nuchal and dorsal crests, distinctly heterogenous scales on the flanks, a gular crest, two large tubercular scales ventral to the tympanum, and 97–125 scales around midbody will distinguish the new species from all congeners. The new species occurs above 1100 m on mountains of Sumatra's southern Bukit Barisan Range below 3°S latitude. Head scales of some specimens were infested by a green, filamentous symbiont, which we tentatively identify as an epizoic plant. Gravid females develop crimson stripes on their gular pouch as an apparent social signal.




Gonocephalus semperi PETERS,1867

Mindoro Forest Dragon

PETERS, W.C.H. (1867): Herpetologische Notizen. - Monatsber. königl. Akad. Wiss. Berlin. 1867: 13-37.



Gonocephalus sophiae GRAY,1845

Negros Forest Dragon
GAULKE, M. & A. DEMEGILLO (2005): Zur Variabilität und Biologie der philippinischen Winkelkopfagame Gonocephalus sophiae. – Reptilia, Münster, 10 (1): 32-40.

Inhalt:
Beschreibung, Lebensraum, Gelegedaten und Inkubationszeit, Nachwuchs, Ernährung, Verhalten, Danksagung.

GAULKE, M. & A. DEMEGILLO (2005): Biology and variability of the Negros Forest Dragon Gonocephalus sophiae. - Reptilia (GB) (42): 16-23.

GRAY, J.E. (1845): Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp.

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