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Literatur und Schriften
Australische Bodenagamen The Australian dragon lizard genus Diporiphora currently comprises 21 species based on genetic and morphological evidence, with 11 of these species occurring in the monsoonal tropics of northern Australia. Diporiphora are climbing lizards that are found on either trees, grasses or rocks, with usually only subtle morphological differences to distinguish between species. Since the last taxonomic treatment of this genus in northern Australia over 40 years ago, species delimitation using genetic techniques has clarified the number of lineages and increased collections from recent surveys have significantly broadened the distributions of these taxa. However, no formal taxonomic assessments have been undertaken to redefine species, including the many lineages that represent undescribed species. Currently, there are seven species of Diporiphora with vast distributions across northern Australia and a broad and variable set of morphological characteristics that make species identification challenging, even for experienced field workers. Here, we provide a comprehensive taxonomic treatment of Diporiphora species across northern Australia based on previously published genetic data and morphological examination of voucher specimens. Our analyses demonstrate that these broadly distributed taxa actually comprise multiple, often allopatric, species, with especially high diversity in the Kimberley region of Western Australia. We redescribe nine previously described species and describe five new species of Diporiphora based on historical types, newly collected material and older museum vouchers. In the D. australis species group, we resurrect D. jugularis Macleay from synonomy. In the D. bennettii species group, we synonomise D. arnhemica Storr with D. albilabris Storr, and raise to full species the latter and D. sobria Storr. In addition, we describe as new a wide-ranging saxicoline species previously attributed to D. bennettii Gray. In the D. bilineata species group, we resurrect D. margaretae Storr from synonomy with D. magna Storr and describe three new species. Lastly, we describe a species from the northwest Kimberley that is more closely related to an arid zone radiation. The revision of the northern Diporiphora dragons here stabilises the taxonomy, redefines many species distributions and reveals many new species. Further work on Diporiphora includes further surveys to better understand distributions and habitat preferences and continue to refine their evolutionary history and biogeography in northern Australia.
Carnarvon Dragon DOUGHTY, P., KEALLEY, L. & J. MELVILLE (2012): Taxonomic assessment of Diporiphora (Reptilia: Agamidae) dragon lizards from the western arid zone of Australia. – Zootaxa, 3518: 1-24. Members of the genus Diporiphora are slender perching agamid lizards from Australasia, with a conservative morphology and some outstanding taxonomic issues. Here we assess morphological variation in the morphologically similar D. pin-dan, D. valens, and D. winneckei from the western deserts of Australia. A reassessment of morphological differences that included the presence or absence of a gular fold, revealed D. pindan to be much more widely distributed than previously thought, occurring as far south as the northern Pilbara and east to the Tanami Desert. Examination of D. valens specimens revealed a north-south split within the Pilbara, with specimens conspecific with the types from the Hamersley Range in the southern Pilbara, whereas recently collected specimens from the Chichester and Roebourne regions in the northern Pilbara differ morphologically, and are described as a new species. Examination of the type of D. winneckei and topotypic material indicates that populations referable to this species are confined to the eastern arid zone. The isolated far western population of ‘D. winneckei’ from the Carnarvon Basin differs in morphology from the eastern arid zone D. winneckei and is described as a new species. The western arid zone ‘D. winneckei’ is also morphologically distinctive from the eastern arid zone D. winneckei and is described as a new species. We also redescribe D. pindan, D. valens, and D. winneckei, and return Caimanops amphiboluroides to Diporiphora based on the results of previous genetic studies.
White-lipped Two-line Dragon STORR, G.M. (1974): Agamid lizards of the genera Caimanops, Physignathus and Diporiphora in Western Australia and Northern Territory. – Records W. Aust. Mus., 3 (2): 121-146. Caimanopsgen. novo is proposed for Diporiphora amphiboluroides Lucas & Frost. The following species and subspecies of Physignathus and Diporiphora are studied: P. longirostris (Boulenger), P. temporalis (Giinther), P. g. gilberti (Gray), P. g. centralis Loveridge, D. convergens nov., D. a. albilabris nov., D. a. sobria nov., D. b. bennettii (GraY), D. b. arnhemica nov., D. magna nov., D. lalliae nov., D. reginae Glauert, D. winneckei Lucas & Frost, D. b. bilineata Gray, D. b. margaretae nov., and D. superba novo
COUPER, P., MELVILLE, J., EMMOTT, A. & S.N.J. CHAPPLE (2012): A new species of Diporiphora from the Goneaway Tablelands of Western Queensland. Zootaxa 3556: 39–54. Australia’s agamid genus Diporiphora is speciose and widespread, however, there remain significant taxonomic uncertainties within this group. Field collections across the range of Diporiphora continue to uncover undocumented morphological and ecological variation. A new morpho-type was collected from hard pebbly soils on Valetta Station, western Queensland, providing ample data for the description of a new species (Diporiphora ameliae sp. nov.). We undertook a morphological study, integrated with a comprehensive genetic study to provide the phylogenetic placement and distinctiveness of the new species. Although superficially similar to Diporiphora winneckei, the new species is characterised by well developed ventral body patterns consisting of four longitudinal grey stripes on a cream background and three distinctive dark V-shaped markings that converge anteriorly on the throat and gular area. Molecular data is presented incorporating a ~1200 bp of the mtDNA protein-coding gene ND2 and five flanking tRNAs for 58 new sequences and 53 previously published sequences. Phylogenetic analyses of the molecular data strongly support the new species as an independent evolutionary lineage within Diporiphora. In addition, the molecular data also showed that there is far greater diversity in Diporiphora winneckei sensu lato than was anticipated. Our results clearly indicate that there are at least three independent evolutionary lineages of D. winneckei-like dragons.
GOLDBERG, S.R. (2009): Notes on reproduction of the Mulga dragon, Caimanops amphiboluroides, and the western netted dragon, Ctenophorus reticulatus, (Squamata: Agamidae) from Western Australia. – Bull. Chic. Herp. Soc., 44 (7): 114-115.
Mulga Dragon Histological analyses were conducted on gonadal material of two Australian agamid lizards, Caimanops amphiboluroides and Ctenophorus reticulatus. Caimanops amphiboluroides males from December and February were undergoing spermiogenesis. Females of C. amphiboluroides from December contained eight and ten enlarged follicles (> 4 mm), respectively. Ctenophorus reticulatus males from October, December, January and May were undergoing spermiogenesis. One female C. reticulatus from October contained corpora lutea from a previous egg clutch and concomitant yolk deposition for a subsequent clutch indicating production of multiple clutches in the same reproductive season. It appears the reproductive cycles of C. amphiboluroides and C. reticulatus most closely fit the Type I category of Heatwole and Taylor (1987) in which spermatogenesis and ovulation occur in spring. Ecological data on the seldom encountered and little known agamid Caimanops amphiboluroides are presented. Caimanops is arboreal and is usually associated with mulga trees and fence posts. Average height above ground of 10 active lizards was 79.5 cm. Average body temperature of 13 active lizards was 36.6°C. These lizards are dietary specialists, feeding mostly on termites. Average clutch size is 12 (N=2). Caimanopsgen. novo is proposed for Diporiphora amphiboluroides Lucas & Frost. The following species and subspecies of Physignathus and Diporiphora are studied: P. longirostris (Boulenger), P. temporalis (Giinther), P. g. gilberti (Gray), P. g. centralis Loveridge, D. convergens nov., D. a. albilabris nov., D. a. sobria nov., D. b. bennettii (GraY), D. b. arnhemica nov., D. magna nov., D. lalliae nov., D. reginae Glauert, D. winneckei Lucas & Frost, D. b. bilineata Gray, D. b. margaretae nov., and D. superba novo.
Eastern two-line Dragon DE VIS, C.W. (1884): On new species of Australian lizards. – Proceedings of the Ropyal Society of Queensland, 1: 97-100. EDWARDS, D.L. & J. MELVILLE (2010): Phylogeographic analysis detects congruent biogeographic patterns between a woodland agamid and Australian wet tropics taxa despite disparate evolutionary trajectories. – J. Biogeogr., 37 (8): 1543-1556. Aim To test the congruence of phylogeographic patterns and processes between a woodland agamid lizard (Diporiphora australis) and well-studied Australian wet tropics fauna. Specifically, to determine whether the biogeographic history of D. australis is more consistent with a history of vicariance, which is common in wet tropics fauna, or with a history of dispersal with expansion, which would be expected for species occupying woodland habitats that expanded with the increasingly drier conditions in eastern Australia during the Miocene–Pleistocene. Location North-eastern Australia. Methods Field-collected and museum tissue samples from across the entire distribution of D. australis were used to compile a comprehensive phylogeographic dataset based on c. 1400 bp of mitochondrial DNA (mtDNA), incorporating the ND2 protein-coding gene. We used phylogenetic methods to assess biogeographic patterns within D. australis and relaxed molecular clock analyses were conducted to estimate divergence times. Hierarchical Shimodaira– Hasegawa tests were used to test alternative topologies representing vicariant, dispersal and mixed dispersal/vicariant biogeographic hypotheses. Phylogenetic analyses were combined with phylogeographic analyses to gain an insight into the evolutionary processes operating within D. australis. Results Phylogenetic analyses identified six major mtDNA clades within D. australis, with phylogeographic patterns closely matching those seen in many wet tropics taxa. Congruent phylogeographic breaks were observed across the Black Mountain Corridor, Burdekin and St Lawrence Gaps. Divergence amongst clades was found to decrease in a north–south direction, with a trend of increasing population expansion in the south. Main conclusions While phylogeographic patterns in D. australis reflect those seen in many rain forest fauna of the wet tropics, the evolutionary processes underlying these patterns appear to be very different. Our results support a history of sequential colonization of D. australis from north to south across major biogeographic barriers from the late Miocene–Pleistocene. These patterns are most likely in response to expanding woodland habitats. Our results strengthen the data available for this iconic region in Australia by exploring the understudied woodland habitats. In addition, our study shows the importance of thorough investigations of not only the biogeographic patterns displayed by species but also the evolutionary processes underlying such patterns
Robust Two-line Dragon GRAY, J.E. (1845): Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp.
Northern Two-line Dragon, Two-Lined Dragon GRAY, J.E. (1842): Description of some hitherto unrecorded species of Australian reptiles and batrachians.- Zoological Miscellany 2: 51-57 (London: Treuttel, Würtz & Co). MARYAN, B. (1995): A note on colour change in Diporiphora bilineata. – Herpetofauna, Sydney, 25 (2): 61.
Crystal Creek Two-line Dragon STORR, G.M. (1974): Agamid lizards of the genera Caimanops, Physignathus and Diporiphora in Western Australia and Northern Territory. – Records W. Aust. Mus., 3 (2): 121-146. Caimanopsgen. novo is proposed for Diporiphora amphiboluroides Lucas & Frost. The following species and subspecies of Physignathus and Diporiphora are studied: P. longirostris (Boulenger), P. temporalis (Giinther), P. g. gilberti (Gray), P. g. centralis Loveridge, D. convergens nov., D. a. albilabris nov., D. a. sobria nov., D. b. bennettii (GraY), D. b. arnhemica nov., D. magna nov., D. lalliae nov., D. reginae Glauert, D. winneckei Lucas & Frost, D. b. bilineata Gray, D. b. margaretae nov., and D. superba novo.
Gracile Two-lined Dragon MELVILLE, J., SMITH DATE, K.L., HORNER, P. & P. DOUGHTY (2019): Taxonomic revision of dragon lizards in the genus Diporiphora (Reptilia: Agamidae) from the Australian monsoonal tropics. Memoirs of Museum Victoria 78: 23–55. The Australian dragon lizard genus Diporiphora currently comprises 21 species based on genetic and morphological evidence, with 11 of these species occurring in the monsoonal tropics of northern Australia. Diporiphora are climbing lizards that are found on either trees, grasses or rocks, with usually only subtle morphological differences to distinguish between species. Since the last taxonomic treatment of this genus in northern Australia over 40 years ago, species delimitation using genetic techniques has clarified the number of lineages and increased collections from recent surveys have significantly broadened the distributions of these taxa. However, no formal taxonomic assessments have been undertaken to redefine species, including the many lineages that represent undescribed species. Currently, there are seven species of Diporiphora with vast distributions across northern Australia and a broad and variable set of morphological characteristics that make species identification challenging, even for experienced field workers. Here, we provide a comprehensive taxonomic treatment of Diporiphora species across northern Australia based on previously published genetic data and morphological examination of voucher specimens. Our analyses demonstrate that these broadly distributed taxa actually comprise multiple, often allopatric, species, with especially high diversity in the Kimberley region of Western Australia. We redescribe nine previously described species and describe five new species of Diporiphora based on historical types, newly collected material and older museum vouchers. In the D. australis species group, we resurrect D. jugularis Macleay from synonomy. In the D. bennettii species group, we synonomise D. arnhemica Storr with D. albilabris Storr, and raise to full species the latter and D. sobria Storr. In addition, we describe as new a wide-ranging saxicoline species previously attributed to D. bennettii Gray. In the D. bilineata species group, we resurrect D. margaretae Storr from synonomy with D. magna Storr and describe three new species. Lastly, we describe a species from the northwest Kimberley that is more closely related to an arid zone radiation.The revision of the northern Diporiphora dragons here stabilises the taxonomy, redefines many species distributions and reveals many new species. Further work on Diporiphora includes further surveys to better understand distributions and habitat preferences and continue to refine their evolutionary history and biogeography in northern Australia.
Granulated two-lined Dragon MELVILLE, J., SMITH DATE, K.L., HORNER, P. & P. DOUGHTY (2019): Taxonomic revision of dragon lizards in the genus Diporiphora (Reptilia: Agamidae) from the Australian monsoonal tropics. Memoirs of Museum Victoria 78: 23–55. The Australian dragon lizard genus Diporiphora currently comprises 21 species based on genetic and morphological evidence, with 11 of these species occurring in the monsoonal tropics of northern Australia. Diporiphora are climbing lizards that are found on either trees, grasses or rocks, with usually only subtle morphological differences to distinguish between species. Since the last taxonomic treatment of this genus in northern Australia over 40 years ago, species delimitation using genetic techniques has clarified the number of lineages and increased collections from recent surveys have significantly broadened the distributions of these taxa. However, no formal taxonomic assessments have been undertaken to redefine species, including the many lineages that represent undescribed species. Currently, there are seven species of Diporiphora with vast distributions across northern Australia and a broad and variable set of morphological characteristics that make species identification challenging, even for experienced field workers. Here, we provide a comprehensive taxonomic treatment of Diporiphora species across northern Australia based on previously published genetic data and morphological examination of voucher specimens. Our analyses demonstrate that these broadly distributed taxa actually comprise multiple, often allopatric, species, with especially high diversity in the Kimberley region of Western Australia. We redescribe nine previously described species and describe five new species of Diporiphora based on historical types, newly collected material and older museum vouchers. In the D. australis species group, we resurrect D. jugularis Macleay from synonomy. In the D. bennettii species group, we synonomise D. arnhemica Storr with D. albilabris Storr, and raise to full species the latter and D. sobria Storr. In addition, we describe as new a wide-ranging saxicoline species previously attributed to D. bennettii Gray. In the D. bilineata species group, we resurrect D. margaretae Storr from synonomy with D. magna Storr and describe three new species. Lastly, we describe a species from the northwest Kimberley that is more closely related to an arid zone radiation.The revision of the northern Diporiphora dragons here stabilises the taxonomy, redefines many species distributions and reveals many new species. Further work on Diporiphora includes further surveys to better understand distributions and habitat preferences and continue to refine their evolutionary history and biogeography in northern Australia.
Black-throated two-pored dragon MACLEAY, W. (1877): The lizards of the Chevert Expedition. - Proceedings of the Linnean Society of New South Wales, 2: 60-69; 97-104.
Lally´s Two-line Dragon STORR, G.M. (1974): Agamid lizards of the genera Caimanops, Physignathus and Diporiphora in Western Australia and Northern Territory. – Records W. Aust. Mus., 3 (2): 121-146. Caimanopsgen. novo is proposed for Diporiphora amphiboluroides Lucas & Frost. The following species and subspecies of Physignathus and Diporiphora are studied: P. longirostris (Boulenger), P. temporalis (Giinther), P. g. gilberti (Gray), P. g. centralis Loveridge, D. convergens nov., D. a. albilabris nov., D. a. sobria nov., D. b. bennettii (GraY), D. b. arnhemica nov., D. magna nov., D. lalliae nov., D. reginae Glauert, D. winneckei Lucas & Frost, D. b. bilineata Gray, D. b. margaretae nov., and D. superba novo.
Pink Two-line Dragon HOUSTON, T.F. (1977): A new species of Diporiphora from South Australia and geographic variation in D. winneckei Lucas & Frost (Lacertilia: Agamidae). – Transactions R. Soc. S. Aust., 101 (8): 199-205.
Yellow-sided Two-line Dragon STORR, G.M. (1974): Agamid lizards of the genera Caimanops, Physignathus and Diporiphora in Western Australia and Northern Territory. – Records W. Aust. Mus., 3 (2): 121-146. Caimanopsgen. novo is proposed for Diporiphora amphiboluroides Lucas & Frost. The following species and subspecies of Physignathus and Diporiphora are studied: P. longirostris (Boulenger), P. temporalis (Giinther), P. g. gilberti (Gray), P. g. centralis Loveridge, D. convergens nov., D. a. albilabris nov., D. a. sobria nov., D. b. bennettii (GraY), D. b. arnhemica nov., D. magna nov., D. lalliae nov., D. reginae Glauert, D. winneckei Lucas & Frost, D. b. bilineata Gray, D. b. margaretae nov., and D. superba novo.
STORR, G.M. (1974): Agamid lizards of the genera Caimanops, Physignathus and Diporiphora in Western Australia and Northern Territory. – Records W. Aust. Mus., 3 (2): 121-146. Caimanopsgen. novo is proposed for Diporiphora amphiboluroides Lucas & Frost. The following species and subspecies of Physignathus and Diporiphora are studied: P. longirostris (Boulenger), P. temporalis (Giinther), P. g. gilberti (Gray), P. g. centralis Loveridge, D. convergens nov., D. a. albilabris nov., D. a. sobria nov., D. b. bennettii (GraY), D. b. arnhemica nov., D. magna nov., D. lalliae nov., D. reginae Glauert, D. winneckei Lucas & Frost, D. b. bilineata Gray, D. b. margaretae nov., and D. superba novo.
DRISCOLL, D.A. & C.M. HARDY (2005): Dispersal and phylogeography of the agamid lizard Amphibolurus nobbi in fragmented and continuous habitat. – Molec. Ecol., 14 (6): 1613-1629. Approximately 90% of native vegetation has been cleared for agriculture in central New South Wales, Australia. Habitat loss has reduced and fragmented populations of the agamid lizard Amphibolurus nobbi. We compared genetic structure of populations of this species in an unmodified landscape with those from small nature reserves and linear remnants in farming areas. We ask: Is there evidence for reduced dispersal and population fragmentation among farm populations? Using 2008 bp mtDNA sequences and allozyme electrophoresis, we found that small populations in farming areas had as much genetic variation as populations in nature reserves. Application of nested clade analysis (NCA) indicated isolation-by-distance effects among populations from uncleared areas, but not among populations within farming locations. The genetic evidence therefore implied a high level of migration in the cleared landscapes. High dispersal after fragmentation may have resulted from either a burst of movement at the time of land clearing with dragons from many sources finding refuge in a few remnants, or from ongoing rapid dispersal through unsuitable habitat. A phylogeny based on mtDNA revealed that A. nobbi populations in the study area are deeply divided into two reciprocally monophyletic groups. Although we did not sample the entire species range, one of these evolutionarily significant units was only detected in remnant vegetation in the agricultural landscape. Therefore, a substantial subclade of this species may be vulnerable to extinction. Our findings emphasize that local populations of widespread species can harbour important intraspecific genetic diversity, supporting the case for maintaining widespread species throughout production landscapes. Morphological and molecular information is invaluable in the description of cryptic diversity and the evolutionary processes driving diversification within closely related species that exhibit morphological homoplasy. We present a distribution-wide data set consisting of both molecular and morphological information, providing a taxonomic revision of the Diporiphora nobbi species group, and develop preliminary hypotheses regarding the evolutionary history of D. nobbi. We show deep molecular divergence between D. nobbi and a newly described sister lineage associated with divergence in meristic characters. Our molecular data also show large divergences among subclades within nominate D. nobbi associated with different habitats rather than specific biogeographc barriers. We further discuss potential diversification mechanisms within the D. nobbi species group.
Paled two-pored Dragon MELVILLE, J., SMITH DATE, K.L., HORNER, P. & P. DOUGHTY (2019): Taxonomic revision of dragon lizards in the genus Diporiphora (Reptilia: Agamidae) from the Australian monsoonal tropics. Memoirs of Museum Victoria 78: 23–55. The Australian dragon lizard genus Diporiphora currently comprises 21 species based on genetic and morphological evidence, with 11 of these species occurring in the monsoonal tropics of northern Australia. Diporiphora are climbing lizards that are found on either trees, grasses or rocks, with usually only subtle morphological differences to distinguish between species. Since the last taxonomic treatment of this genus in northern Australia over 40 years ago, species delimitation using genetic techniques has clarified the number of lineages and increased collections from recent surveys have significantly broadened the distributions of these taxa. However, no formal taxonomic assessments have been undertaken to redefine species, including the many lineages that represent undescribed species. Currently, there are seven species of Diporiphora with vast distributions across northern Australia and a broad and variable set of morphological characteristics that make species identification challenging, even for experienced field workers. Here, we provide a comprehensive taxonomic treatment of Diporiphora species across northern Australia based on previously published genetic data and morphological examination of voucher specimens. Our analyses demonstrate that these broadly distributed taxa actually comprise multiple, often allopatric, species, with especially high diversity in the Kimberley region of Western Australia. We redescribe nine previously described species and describe five new species of Diporiphora based on historical types, newly collected material and older museum vouchers. In the D. australis species group, we resurrect D. jugularis Macleay from synonomy. In the D. bennettii species group, we synonomise D. arnhemica Storr with D. albilabris Storr, and raise to full species the latter and D. sobria Storr. In addition, we describe as new a wide-ranging saxicoline species previously attributed to D. bennettii Gray. In the D. bilineata species group, we resurrect D. margaretae Storr from synonomy with D. magna Storr and describe three new species. Lastly, we describe a species from the northwest Kimberley that is more closely related to an arid zone radiation.The revision of the northern Diporiphora dragons here stabilises the taxonomy, redefines many species distributions and reveals many new species. Further work on Diporiphora includes further surveys to better understand distributions and habitat preferences and continue to refine their evolutionary history and biogeography in northern Australia
Grey-striped Western Desert Dragon DOUGHTY, P., KEALLEY, L. & J. MELVILLE (2012): Taxonomic assessment of Diporiphora (Reptilia: Agamidae) dragon lizards from the western arid zone of Australia. – Zootaxa, 3518: 1-24. Members of the genus Diporiphora are slender perching agamid lizards from Australasia, with a conservative morphology and some outstanding taxonomic issues. Here we assess morphological variation in the morphologically similar D. pin-dan, D. valens, and D. winneckei from the western deserts of Australia. A reassessment of morphological differences that included the presence or absence of a gular fold, revealed D. pindan to be much more widely distributed than previously thought, occurring as far south as the northern Pilbara and east to the Tanami Desert. Examination of D. valens specimens revealed a north-south split within the Pilbara, with specimens conspecific with the types from the Hamersley Range in the southern Pilbara, whereas recently collected specimens from the Chichester and Roebourne regions in the northern Pilbara differ morphologically, and are described as a new species. Examination of the type of D. winneckei and topotypic material indicates that populations referable to this species are confined to the eastern arid zone. The isolated far western population of ‘D. winneckei’ from the Carnarvon Basin differs in morphology from the eastern arid zone D. winneckei and is described as a new species. The western arid zone ‘D. winneckei’ is also morphologically distinctive from the eastern arid zone D. winneckei and is described as a new species. We also redescribe D. pindan, D. valens, and D. winneckei, and return Caimanops amphiboluroides to Diporiphora based on the results of previous genetic studies.
Kimberley Rock Dragon MELVILLE, J., SMITH DATE, K.L., HORNER, P. & P. DOUGHTY (2019): Taxonomic revision of dragon lizards in the genus Diporiphora (Reptilia: Agamidae) from the Australian monsoonal tropics. Memoirs of Museum Victoria 78: 23–55. The Australian dragon lizard genus Diporiphora currently comprises 21 species based on genetic and morphological evidence, with 11 of these species occurring in the monsoonal tropics of northern Australia. Diporiphora are climbing lizards that are found on either trees, grasses or rocks, with usually only subtle morphological differences to distinguish between species. Since the last taxonomic treatment of this genus in northern Australia over 40 years ago, species delimitation using genetic techniques has clarified the number of lineages and increased collections from recent surveys have significantly broadened the distributions of these taxa. However, no formal taxonomic assessments have been undertaken to redefine species, including the many lineages that represent undescribed species. Currently, there are seven species of Diporiphora with vast distributions across northern Australia and a broad and variable set of morphological characteristics that make species identification challenging, even for experienced field workers. Here, we provide a comprehensive taxonomic treatment of Diporiphora species across northern Australia based on previously published genetic data and morphological examination of voucher specimens. Our analyses demonstrate that these broadly distributed taxa actually comprise multiple, often allopatric, species, with especially high diversity in the Kimberley region of Western Australia. We redescribe nine previously described species and describe five new species of Diporiphora based on historical types, newly collected material and older museum vouchers. In the D. australis species group, we resurrect D. jugularis Macleay from synonomy. In the D. bennettii species group, we synonomise D. arnhemica Storr with D. albilabris Storr, and raise to full species the latter and D. sobria Storr. In addition, we describe as new a wide-ranging saxicoline species previously attributed to D. bennettii Gray. In the D. bilineata species group, we resurrect D. margaretae Storr from synonomy with D. magna Storr and describe three new species. Lastly, we describe a species from the northwest Kimberley that is more closely related to an arid zone radiation.The revision of the northern Diporiphora dragons here stabilises the taxonomy, redefines many species distributions and reveals many new species. Further work on Diporiphora includes further surveys to better understand distributions and habitat preferences and continue to refine their evolutionary history and biogeography in northern Australia
EDWARDS, D.L. & J. MELVILLE (2011): Extensive phylogeographic and morphological diversity in Diporiphora nobbi (Agamidae) leads to a taxonomic review and a new species description. – Journal of Herpetology, 45 (4); 530-546. Morphological and molecular information is invaluable in the description of cryptic diversity and the evolutionary processes driving diversification within closely related species that exhibit morphological homoplasy. We present a distribution-wide data set consisting of both molecular and morphological information, providing a taxonomic revision of the Diporiphora nobbi species group, and develop preliminary hypotheses regarding the evolutionary history of D. nobbi. We show deep molecular divergence between D. nobbi and a newly described sister lineage associated with divergence in meristic characters. Our molecular data also show large divergences among subclades within nominate D. nobbi associated with different habitats rather than specific biogeographc barriers. We further discuss potential diversification mechanisms within the D. nobbi species group.
Pindan Two-line Dragon McALPIN, S. (1996): A range extension for the dragon Diporiphora pindan. – Herpetofauna, Sydney, 26 (1): 47. STORR, G.M. (1980): Two new Diporiphora (Lacertilia, Agamidae) from Western Australia. – Rec. West. Aust. Mus., 7 (2): 255-263.
Plain-backed Two-line Dragon GLAUERT, L. (1959): A new Agamid lizard from Queen Victoria Springs, Western Australia (Diporiphora reginae, sp. nov.). – Proc. R. zool. Soc. N.S.W., 1959: 10.
Northern savannah two-pored dragon STORR, G.M. (1974): Agamid lizards of the genera Caimanops, Physignathus and Diporiphora in Western Australia and Northern Territory. – Records W. Aust. Mus., 3 (2): 121-146. Caimanopsgen. novo is proposed for Diporiphora amphiboluroides Lucas & Frost. The following species and subspecies of Physignathus and Diporiphora are studied: P. longirostris (Boulenger), P. temporalis (Giinther), P. g. gilberti (Gray), P. g. centralis Loveridge, D. convergens nov., D. a. albilabris nov., D. a. sobria nov., D. b. bennettii (GraY), D. b. arnhemica nov., D. magna nov., D. lalliae nov., D. reginae Glauert, D. winneckei Lucas & Frost, D. b. bilineata Gray, D. b. margaretae nov., and D. superba novo.
Superb Two-line Dragon STORR, G.M. (1974): Agamid lizards of the genera Caimanops, Physignathus and Diporiphora in Western Australia and Northern Territory. – Records W. Aust. Mus., 3 (2): 121-146.Caimanopsgen. novo is proposed for Diporiphora amphiboluroides Lucas & Frost. The following species and subspecies of Physignathus and Diporiphora are studied: P. longirostris (Boulenger), P. temporalis (Giinther), P. g. gilberti (Gray), P. g. centralis Loveridge, D. convergens nov., D. a. albilabris nov., D. a. sobria nov., D. b. bennettii (GraY), D. b. arnhemica nov., D. magna nov., D. lalliae nov., D. reginae Glauert, D. winneckei Lucas & Frost, D. b. bilineata Gray, D. b. margaretae nov., and D. superba novo. WEIGEL, J. (1989): Maintenance and breeding of the superb dragon Diporiphora superba at the Australian Reptile Park, Gosford. – Int. Zoo Yb., 28: 122-126.
Pilbara Two-line Dragon MARYAN, B. & J. TURPIN (2012): Natural history notes on two Pilbara endemic lizards, Diporiphora valens and Varanus bushi, with brief comments on survey methodology. - The Western Australian Naturalist, 28: 274-279. STORR, G.M. (1980): Two new Diporiphora (Lacertilia, Agamidae) from Western Australia. – Rec. West. Aust. Mus., 7 (2): 255-263.
Northern Pilbara Tree Dragon DOUGHTY, P., KEALLEY, L. & J. MELVILLE (2012): Taxonomic assessment of Diporiphora (Reptilia: Agamidae) dragon lizards from the western arid zone of Australia. – Zootaxa, 3518: 1-24. Members of the genus Diporiphora are slender perching agamid lizards from Australasia, with a conservative morphology and some outstanding taxonomic issues. Here we assess morphological variation in the morphologically similar D. pin-dan, D. valens, and D. winneckei from the western deserts of Australia. A reassessment of morphological differences that included the presence or absence of a gular fold, revealed D. pindan to be much more widely distributed than previously thought, occurring as far south as the northern Pilbara and east to the Tanami Desert. Examination of D. valens specimens revealed a north-south split within the Pilbara, with specimens conspecific with the types from the Hamersley Range in the southern Pilbara, whereas recently collected specimens from the Chichester and Roebourne regions in the northern Pilbara differ morphologically, and are described as a new species. Examination of the type of D. winneckei and topotypic material indicates that populations referable to this species are confined to the eastern arid zone. The isolated far western population of ‘D. winneckei’ from the Carnarvon Basin differs in morphology from the eastern arid zone D. winneckei and is described as a new species. The western arid zone ‘D. winneckei’ is also morphologically distinctive from the eastern arid zone D. winneckei and is described as a new species. We also redescribe D. pindan, D. valens, and D. winneckei, and return Caimanops amphiboluroides to Diporiphora based on the results of previous genetic studies.
Canegrass Two-line Dragon GOLDBERG, S.R. (2007): Notes on reproduction of the lozenge-marked bicycle-dragon, Ctenophorus scutulatus and the canegrass two-line dragon, Diporiphora winneckei (Squamata: Agamidae) from Western Australia. – Bull. Mary. Herp. Soc., 43 (1): 1-3. HOUSTON, T.F. (1977): A new species of Diporiphora from South Australia and geographic variation in D. winneckei Lucas & Frost (Lacertilia: Agamidae). – Transactions R. Soc. S. Aust., 101 (8): 199-205. LUCAS, A.H S. & C. FROST (1896): Further preliminary notice of certain new species of lizards from central Australia. - Proceedings of the Royal Society of Victoria, 8: 1-4. zurück / back |
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