Literatur und Schriften


Bronchocela KAUP, 1827

Langschwanzagamen

BOBROV, V.V. (2017): Lizards (Reptilia, Sauria) from Vietnam in the herpetological collection of the Zoological Museum of Moscow University. 3. genus Bronchocela (Family Agamidae). – Current Studies in Herpetology, 17 (3/4): 198-202. (In Russisch)

Data are given on the lizard specimens of the genus Bronchocela (the family Agamidae) of the Vietnam fauna, accumulated in the herpetological collection of the Zoological Museum of Moscow University. The following data are presented for each specimen: the accession number, the date and location of capture, the habitat, and the collector. The places of capture are shown on the map of the country. A new key for the Vietnamese species of Bronchocela is provided.

GRISMER, L.L., WOOD JR P.L., LEE, C.H, QUAH, E.S.H., ANUAR, S., NGADI, E. & J.W. SITES, JR. (2015): An integrative taxonomic review of the agamid genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with descriptions of new montane and insular endemics. - Zootaxa 3948 (1): 001-023.

An integrative taxonomic analysis is used to identify and describe two new species of the agamid genus Bronchocela (Kuhl) from Peninsular Malaysia: an upland species B. shenlong sp. nov. from Bukit Larut, Perak in the Bintang Mountain Range and Parit Falls, Cameron Highlands, Pahang in the Titiwangsa Mountain Range and an insular species, B. rayaensis sp. nov., from Pulau Langkawi, Kedah off the northwest coast on the border with Thailand. Both species are diagnosed from each other and all other species of Bronchocela on the basis of body shape, scale morphology, and color pattern. The analysis also demonstrates the remarkable genetic similarity of B. cristatella (Kuhl) throughout 1120 km of its range from northern Peninsular Malaysia to western Borneo despite its highly variable coloration and pattern. The two new species are appended to a rapidly growing list of newly described lizard species (60 to date) from Peninsular Malaysia tallied within the last decade.

HALLERMANN, J. (2003): Biogeography and taxonomy of the genus Bronchocela in SE. Asia, with special reference to the Philippine Islands (Squamata: Agamidae). - In: Ananjeva, N. & O. Tsinenko (Eds.): Herpetologia Petropolitana. Programme and abstracts. 12th ordinary general meeting. Societas Europaea Herpetologica (SEH). Zoological Institute of the Russian Academy of Sciences, Saint Petersburg, Russia, 12-16 August 2003. Societas Europaea Herpetologica (SEH): 67.

HALLERMANN, J. (2005): A taxonomic review of the genus Bronchocela (Squamata: Agamidae), with description of a new species from Vietnam. – Russian Journal of Herpetology, 12 (3):167-182.

HALLERMANN, J. (2009): The genera Pseudocalotes and Bronchocela in south East Asia: Taxonomy and distribution. - Abstracts of presentations hold on DeAGAMIS the 1st International Symposium on Agamid Lizards. Bonner Zoologische Beiträge, Bonn, 56 (4): 301.

KAUP, J. (1827): Description of the genus Bronchocela. – In: “Zoologische Monographien”. - Isis von Oken 20: 610-625.

MANTHEY, U. & W. GROSSMANN (1979): Genus Bronchocela - In: Amphibien und Reptilien Südostasiens. - Natur und Tier-Verlag, Münster: 159-162.


Bronchocela burmana (BLANFORD, 1878)

Burma-Langschwanzagame / Burma-Schönechse

BLANFORD, W.T. (1878): Notes on some Reptilia from the Himalayas and Burma. - J. Asiat. Soc. Bengal (2) xlvii: 125-131.

MULCAHY, D.G., COTA, M., MAKCHAI, S. & B.L. STUART (2021): Molecular and morphological evidence for a significant range extension of Bronchocela burmana Blanford, 1878 to eastern Thailand. - Herpetology Notes, 14: 485-491.

ZUG, G.R., MULCAHY, D.G. & J.V. VINDUM (2017): Resurrection of Bronchocela burmana Blanford, 1878 for the Green Crested Lizard (Squamata, Agamidae) of southern Myanmar. ZooKeys 657: 141-156.

Recent fieldwork in southern Tanintharyi revealed the presence of a small Green Crested Lizard in the wet evergreen forest. We generated mtDNA sequence data (ND2) that demonstrates that this population’s nearest relative is Bronchocela rayaensis Grismer et al., 2015 of Pulau Langkawi, northwestern Peninsular Malaysia and Phuket Island. Morphologically the Burmese Bronchocela shares many features with B. rayaensis, which potentially would make this recently described Thai-Malay species a synonym of Bronchocela burmana Blanford, 1878; however, we interpret the genetic and morphological differences to reflect evolutionary divergence and recommend the recognition of both species.


Bronchocela celebensis GRAY, 1845

Sulawesi Bloodsucker



Bronchocela cristatella (KUHL, 1820)

Borneo-Langschwanzagame / Borneo-Schönechse / Blutsaugeragame

BAEDERMANN, L. (2007): Borneo-Schönechsen (Bronchocela cristatella) im Terrarium – schwierig und heikel? – Terraria, Münster, 2 (5): 63-67.

BLUMBERG, A. (1983): Das Portrait: Calotes cristatellus (KUHL, 1820) Borneo-Schönechse. – Sauria, Berlin, 5 (4): 1-2.

DAS, I. (2013): Bronchocela cristatella (crested green lizard) attempted prey. -  Herp. Rev. 44 (2): 319-320.


DAS, I. & R. GEMEL (2000): Nomenclatural status of Fitzinger’s (1861) Pseudocalotes archiducissae, and confirmation of Bronchocela cristatella (Kuhl, 1820) from the Nicobar Archipelago (Squamata: sauri: Agamidae). – Herpetozoa, 13 (1/2): 55-58.

DIONG, C.H. & S.S.L. LIM (1998): Taxonomic review and morphological description of Bronchocela cristatella (Kuhl, 1820) (Squamata: Agamidae) with notes on other species in the genus. – Raffles Bulletin of Zoology, 46: 345-359.

EVAN, S.H., QUAH, C.T.C., MOHD, A.M. & A.M.S. SHAHRUL (2012): Color changes during courtship and mating of the green crested lizard Bronchocela cristatella (Kuhl, 1820) (Squamata: Agamidae) with a discussion of its behavioral and evolutionary significance. - Russian Journal of Herpetology 19: 303–306.

FITZSIMONS, J. & J. THOMAS (2016): Bronchocela cristatella (Green Crested Lizard). Diet and foraging behavior. - Herp. Rev. 47 (3): 462.

GOLDBERG, S.R. & L.L. GRISMER (2014): Bronchocela cristatella (Green Crested Lizard) – reproduction. - Herp. Rev. 45 (4): 695.

GOLDBERG, S.R. & L.L. GRISMER (2014): Natural history notes: Bronchocela cristatella (Green Crested Lizard). Reproduction. – Herp. Rev., 45 (2): 326-327.

KUHL, H. (1820): Description of Bronchocela cristatella  in “Beiträge zur Zoologie und vergleichenden Anatomie”. Hermannsche Buchhandlung, Frankfurt, 152 pp.

LIAT, L.B. & H.B. PERAL (1959): Paradise tree snake versus green crested lizard. – Malay. nat. J., 14: 33-34.

MANTHEY, U. et al. (1994): Gleitfliegen bei Bronchocela cristatella (KUHL, 1820) (Sauria: Agamidae). - Sauria, Berlin, 16 (3): 37-38. (1223)

RANDOW, H. (1952): Calotes versicolor Daudin und Calotes cristatellus Kühl. – Die Aquar. Terrar. Z., Stuttgart, 5: 185-187.

REID, J.A. (1958): Gliding by the Green Crested lizard (Calotes? cristatellus). – Malay. Nat. J., 12: 119.

YEO, R.K.H. (2013): Green crested lizards mating. -  Singapore Biodiversity Records 2013: 30-31.



Bronchocela cyanopalpebra CHANDRAMOULI, ADHIKARI, AMARASINGHE & ABINAWANTO, 2023

CHANDRAMOULI, S., ADHIKARI, O. D., AMARASINGHE, A. T., & ABINAWANTO, A. 2023. A review of the genus Bronchocela Kaup, 1827 (Reptilia: Agamidae) in the Nicobar Archipelago with the description of two new species. Zootaxa 5254 (4): 493-516



Bronchocela danieli TIWARI & BISWAS, 1973

Daniel´s Forest Lizard

TIWARI, K.K. & S. BISWAS (1973): Two new reptiles from the great Nicobar Islands. - J. Zool. Soc. India, 25: 57-63.



Bronchocela hayeki MÜLLER, 1928

Sumatra Bloodsucker

MÜLLER, L. (1928): Herpetologische Mitteilungen II. Ein neuer Calotes von Sumatra. – Zool. Anz., 77: 67-69.



Bronchocela jubata DUMÉRIL & BIBRON, 1837

Maned Forest Lizard



Bronchocela marmorata GRAY, 1845

Marbled Bloodsucker

GROSMANN, W. (2009): Der Farb- und Zeichnungswechsel bei einer Marmorierten Langschwanzagame, Bronchocela marmorata GRAY, 1845. – Sauria, Berlin, 31 (1): 23-27.

The changes in colour and pattern of a stressed Marbled Slender Agama male along a timeline of five min utes are documented. Kept in an enclosure at a reptile dealer’s at Daet, province Norte, Philippiners, the agama was approached for photographing. Thus stressed, is altered its appearance drastically and rapidly from abright green, through a variety of changing colourations and patterns, to reddish brown with a dark brown pattern on the body and white labials and chin/gular area. These changes are shown in a sequential series of photographs.

KUZMIN, Y., TKACH, V.V. & S.E. BUSH (2012): A new species of Rhabdias (Nematoda: Rhabdiasidae) from Agamid lizards on Luzon Island, Philippines. – Journal of Parasitology, 98 (3): 608-611.

Rhabdias odilebaini n. sp. is described on the basis of specimens found in the lungs of 2 species of agamid lizards: the Philippine flying lizard Draco spilopterus and the marbled bloodsucker Bronchocela marmorata. Specimens were collected in Aurora Province, Luzon Island, Philippines. The new species of Rhabdias is characterized by presence of 4 submedian lips, inconspicuous lateral lips, rounded cross-shaped oral opening, and tail end bent dorsally. This species is morphologically distinct from other Rhabdias spp. that parasitize reptilian and amphibian hosts, including 3 other species known to parasitize lizards of the Agamidae.


Bronchocela nicobarica CHANDRAMOULI, ADHIKARI, AMARASINGHE & ABINAWANTO, 2023

CHANDRAMOULI, S., ADHIKARI, O. D., AMARASINGHE, A. T., & ABINAWANTO, A. 2023. A review of the genus Bronchocela Kaup, 1827 (Reptilia: Agamidae) in the Nicobar Archipelago with the description of two new species. Zootaxa 5254 (4): 493-516



Bronchocela orlovi HALLERMANN, 2004

Orlov´s Forest Lizard

HALLERMANN, J. (2004): A new species of the genus Bronchocela from the tropical rain forest of southern Vietnam. – Russ. J. Herpetol., 11 (1): 30-34.



Bronchocela rayaensis GRISMER, WOOD; LEE, QUAH, ANUAR, NGADI & SITES, 2015

Gunung Raya Green-crested Lizard

GRISMER, L.L., WOOD Jr., P.L., LEE, C.H., QUAH, E.S.H., ANUAR, S., NGADI, E. & J.W. SITES Jr. (2015): An integrative taxonomic review of the agamid genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with descriptions of new montane and insular endemics. Zootaxa, 3948 (1): 001–023.

An integrative taxonomic analysis is used to identify and describe two new species of the agamid genus Bronchocela (Kuhl) from Peninsular Malaysia: an upland species B. shenlong sp. nov. from Bukit Larut, Perak in the Bintang Mountain Range and Parit Falls, Cameron Highlands, Pahang in the Titiwangsa Mountain Range and an insular species, B. rayaensis sp. nov., from Pulau Langkawi, Kedah off the northwest coast on the border with Thailand. Both species are diagnosed from each other and all other species of Bronchocela on the basis of body shape, scale morphology, and color pattern. The analysis also demonstrates the remarkable genetic similarity of B. cristatella (Kuhl) throughout 1120 km of its range from northern Peninsular Malaysia to western Borneo despite its highly variable coloration and pattern. The two new species are appended to a rapidly growing list of newly described lizard species (60 to date) from Peninsular Malaysia tallied with-in the last decade.


Bronchocela rubrigularis HALLERMANN, 2009

HALLERMANN, J. (2009): A new species of Bronchocela (Squamata: Agamidae) from Nicobar Island. – Bonner zoologische Beiträge, Bonn, 56 (4): 279-284.

HALLERMANN, J. (2010): Zwei neue Agamenarten in den Gattungen Pseudocalotes und Bronchocela. – Terraria, Münster, 5 (23): 76-77.



Bronchocela shenlong GRISMER, WOOD, LEE, QUAH, ANUAR, NGADI & SITES, 2015

Diong, Cheong Hoong, and S. S. L. Lim 1998. Taxonomic Review and Morphological Description of Bronchocela cristatella (Kuhl, 1820) (Squamata: Agamidae) with Notes on Other Species in the Genus. Raffles Bulletin of Zoology 46: 345-359.

GOLDBERG, S.R. & L.L. GRISMER (2017): Bronchocela shenlong Reproduction. - Herpetological Review, 48 (1): 186-187.

GOLDBERG, S.R. & L.L. GRISMER (2017): Bronchocela shenlong Endoparasites. - Herpetological Review, 48 (1): 187.

Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155]

Grismer, L.L.; Chan K. Onn, Jesse L. Grismer, Perry L. Wood, Jr., and A. Norhayati 2010. A CHECKLIST OF THE HERPETOFAUNA OF THE BANJARAN BINTANG, PENINSULAR MALAYSIA. Russian Journal of Herpetology 17 (2): 147-160.

GRISMER, L.L.; P.L. WOOD, JR., CHEOL HAENG LEE, EVAN S. H. QUAH, SHAHRUL ANUAR, EHWAN NGADI & JACK 2015. An integrative taxonomic review of the agamid genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with descriptions of new montane and insular endemics. Zootaxa 3948 (1): 001–023.

GRISMER, L.L., WOOD Jr., P.L., LEE, C.H., QUAH, E.S.H., ANUAR, S., NGADI, E. & J.W. SITES Jr. (2015): An integrative taxonomic review of the agamid genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with descriptions of new montane and insular endemics. Zootaxa, 3948 (1): 001–023.

An integrative taxonomic analysis is used to identify and describe two new species of the agamid genus Bronchocela (Kuhl) from Peninsular Malaysia: an upland species B. shenlong sp. nov. from Bukit Larut, Perak in the Bintang Mountain Range and Parit Falls, Cameron Highlands, Pahang in the Titiwangsa Mountain Range and an insular species, B. rayaensis sp. nov., from Pulau Langkawi, Kedah off the northwest coast on the border with Thailand. Both species are diagnosed from each other and all other species of Bronchocela on the basis of body shape, scale morphology, and color pattern. The analysis also demonstrates the remarkable genetic similarity of B. cristatella (Kuhl) throughout 1120 km of its range from northern Peninsular Malaysia to western Borneo despite its highly variable coloration and pattern. The two new species are appended to a rapidly growing list of newly described lizard species (60 to date) from Peninsular Malaysia tallied with-in the last decade.


Bronchocela smaragdina GÜNTHER, 1864

Gunther´s Bloodsucker

GÜNTHER, A. (1864): The Reptiles of British India. - R. Hardwicke, London. i-xxvii + 1-452;  Pl. I-XXVI.



Bronchocela vietnamensis HALLERMANN, 2005

Langschwanzagame

HALLERMANN, J. (2005): A taxonomic review of the genus Bronchocela (Squamata: Agamidae), with description of a new species from Vietnam. – Russian Journal of Herpetology, 12 (3):167-182.

KWET, A. (2006): Bronchocela vietnamensis – eine neue Langschwanzagame aus Vietnam. – Terraria, Münster, 2 (1): 38.

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